Definition: Organisms that manufacture their own organic food from inorganic sources in the non-living environment (autotrophs).
Nearly all utilize photosynthesis; a few use chemosynthesis.
Two Cell Types
Prokaryotic – e.g., cyanobacteria.
Eukaryotic – e.g., algae, plants.
Ecological role: Form the base of food webs; support organisms from zooplankton to large fish (e.g., tuna, sharks).
\textbf{Photoautotroph} – Light energy, \text{CO}_2 carbon source.
Cyanobacteria, plants, algae.
\textbf{Chemoautotroph} – Inorganic chemicals (e.g., \text{NH}3, \text{H}2S), \text{CO}_2 carbon source.
Unique to some bacteria & archaea (e.g., Sulfolobus).
\textbf{Photoheterotroph} – Light energy, organic carbon.
Certain aquatic & halophilic prokaryotes (Rhodobacter, Chloroflexus).
\textbf{Chemoheterotroph} – Organic energy & carbon.
Most prokaryotes, fungi, animals, many protists.
Gram-negative photoautotrophs; first major oxygen producers (>3.5 billion-year-old fossils – oldest known).
Possess internal thylakoid-like membranes; perform \text{N}_2-fixation.
Major component of marine & freshwater phytoplankton; base of aquatic food webs.
Eukaryotes that are not plants, animals, or fungi; most unicellular.
Algae: Aquatic, photosynthetic protists (uni- & multicellular); together with cyanobacteria form primary production in water.
Current hypothesis groups all eukaryotes into four supergroups.
Diatoms – Unicellular algae, glass-like silica wall; highly diverse phytoplankton.
Dinoflagellates – Two flagella, cellulose plates; phototrophs, mixotrophs, heterotrophs; blooms cause toxic “red tides.”
Red Algae – Accessory pigment phycoerythrin masks chlorophyll; mostly multicellular seaweeds; abundant in tropical coasts.
Green Algae – Grass-green chloroplasts; closest relatives of land plants.
Euglenozoa – Spiral/crystalline rod in flagella; mixotrophs, parasites.
Trypanosoma spp. cause Chagas disease & African sleeping sickness.
Lichens: Mutualism between algae (photosynthesis) & fungi (structure); pioneers on bare rock/soil.
Coral Reefs: Algae live in coral polyps, supply food & color; essential for reef building.
Declining protist biomass as sea-surface temperature rises – warm water blocks nutrient upwelling.
Human fertilizer runoff + warming ➔ algal blooms ➔ decomposition creates oxygen-poor dead zones.
Alternation of generations.
Multicellular, dependent embryos.
Apical meristems (localized growth regions).
Mosses, liverworts, hornworts; gametophyte dominant.
Lack true roots (have rhizoids), leaves (have thalli), vascular tissue.
Need water for flagellated sperm.
Sphagnum (peat moss): Forms peat – partially decayed organic matter; fuel source; acidic, low-O₂ peatlands preserve organisms.
Ferns, horsetails; first tall plants (Devonian–Carboniferous).
Vascular tissue allows height:
Xylem – Water & mineral transport; lignified tracheids for support.
Phloem – Distributes sugars & organics.
Still require water for fertilization; formed vast coal forests (Carboniferous).
Origin \approx 360 MYA; dominate terrestrial ecosystems.
Key innovations: reduced gametophytes, heterospory, ovules, pollen, seeds.
Advantages of seeds vs spores: dormancy until favorable, food supply, long-distance dispersal.
4 phyla: Ginkgophyta, Cycadophyta, Gnetophyta, Coniferophyta (largest; evergreen, woody cones).
Medicinal / toxic notes: Ginkgo biloba extracts (memory claims, side effects); Ephedra (ephedrine – stimulant, banned in US).
Reproductive structures: flowers & fruits; phylum Anthophyta.
Flower parts: sepals, petals, stamens (anther + filament), carpels/pistils (stigma, style, ovary).
Double fertilization ➔ seed inside ripened ovary (fruit).
Fruits aid seed dispersal; ethylene (\text{C}2\text{H}4) gas promotes ripening (one bad apple…).
Provide \ge 80\% of human calories – six staples (wheat, rice, maize, potato, cassava, sweet potato).
Plant-derived medicines: Atropine, Digitalin, Menthol, Quinine, Taxol, Tubocurarine, Vinblastine.
Threats: Habitat loss may cause 50\% species extinction within centuries; loss cascades to dependent animals; undiscovered drugs at risk.
Functions: anchor, absorb water/minerals, store carbs.
Root hairs ↑ surface area; mycorrhizal associations.
Modifications:
Storage roots (sweet potato), prop roots (corn), pneumatophores (mangroves for \text{O}_2), buttress roots (tropics), strangling aerial roots (strangler figs).
Nodes (leaf attachment) & internodes.
Buds: Apical (primary elongation) vs Axillary (lateral branches, flowers, thorns).
Modified stems: stolons/runners (asexual spread), rhizomes, tubers (potatoes – eyes = axillary buds), corms, thorns (defense).
Photosynthetic organs; some modifications:
Storage “bulbs” (onion), tendrils (grasp), spines (cacti defense).
Apical meristems: Primary (length) growth.
Lateral meristems: Secondary (thickness) growth.
Vascular cambium – Adds secondary xylem (wood) & phloem.
Cork cambium – Replaces epidermis with protective periderm.
Dendrochronology: Tree-ring analysis for past climate.
Soil texture: sand > silt > clay.
Topsoil: mineral particles + living organisms + humus (decaying organic matter).
Loams: \approx equal sand, silt, clay – most fertile.
Agriculture depletes minerals, water, increases erosion (e.g., Dust Bowl); \sim30\% farmland degraded.
Organic fertilizers (manure, compost) restore nutrients.
Soil organisms decompose organic matter, mix soil.
Crop rotation: Alternate legumes with non-legumes to exploit symbiotic \text{N}_2-fixation.
Special plants: Epiphytes (grow on other plants), parasitic plants (tap host sugars/minerals), carnivorous plants (trap insects to obtain nitrogen).
Multicellular heterotrophs; ingest food.
Unique cell types: nerve & muscle; cells form tissues (functional units).
Many have body cavities (coeloms) – fluid cushions organs, allows independent movement.
Radial (e.g., cnidarians) vs Bilateral (dorsal/ventral, anterior/posterior).
Porifera (Sponges) – No true tissues, filter feeders, sequential hermaphrodites.
Cnidaria – Radial, gastrovascular cavity, stinging cnidocytes; polyp & medusa forms; nerve net; coral reefs build calcium exoskeletons.
Platyhelminthes (Flatworms) – Acoelomate; centralized nervous system.
Free-living planarians regenerate with memory retention.
Parasites: flukes (snail intermediate), tapeworms (no gut; absorb nutrients; pork, beef, flea life cycles).
Nematoda (Roundworms) – Pseudocoelomate; complete gut; no circulatory system; some parasites (hookworm \sim0.5 billion infections, Ascaris \sim1 billion; Trichinella from undercooked pork).
Mollusca – Soft body, CaCO₃ shell (most); open circulatory except cephalopods.
Classes:
• Polyplacophora (chitons)
• Gastropoda (snails, slugs) – many hermaphrodites.
• Bivalvia (clams, oysters, mussels, scallops) – two-part shell, adductor muscles.
• Cephalopoda (squid, octopus) – closed circulation, beak, tentacles, ink, complex brain.
Molluscs suffer highest recorded extinctions.
Annelida – Segmented worms; closed circulation; earthworms (soil aeration, hermaphrodites, can reproduce by fragmentation), leeches (blood-sucking, predators).
Arthropoda – Segmented body, jointed appendages, exoskeleton (chitin), open hemolymph system; >1 million species.
Major lineages: Chelicerates (spiders, ticks), Myriapods (centi- & millipedes), Pancrustaceans (insects, crustaceans).
Insects (Hexapoda): 3 body segments, 3 leg pairs, most undergo metamorphosis (larva \rightarrow pupa \rightarrow adult); internal fertilization; wings in many.
Crustaceans: Isopods, decapods (lobsters, crabs), copepods (zooplankton).
Arachnids: 4 walking leg pairs, pedipalps, chelicerae; many spin silk.
Notochord (flexible rod). 2. Dorsal hollow nerve cord. 3. Pharyngeal slits/clefts. 4. Post-anal tail.
Chondrichthyans – Cartilaginous skeleton (sharks, rays).
Must swim to avoid sinking & ventilate gills.
Acute senses; electrical field detection.
Reproduction:
• \text{Oviparous} – eggs laid.
• \text{Ovoviviparous} – eggs hatch internally.
• \text{Viviparous} – placenta-like nourishment.
Osteichthyes – Bony fish & lobe-fins.
Swim bladder for buoyancy; bony scales; mostly oviparous.
Four limbs with digits, neck, fused pelvic girdle, loss of gills (most), ears.
Frogs, salamanders, caecilians.
Aquatic larva with gills → metamorphosis → terrestrial adult with lungs.
External fertilization; eggs need moisture.
Amniotic egg – Protective shell + extra-embryonic membranes (amnion, chorion, yolk sac, allantois); enables full terrestrial life; rib ventilation, waterproof skin.
Scales (keratin), internal fertilization, shelled eggs, ectothermic (except birds).
Groups: Turtles (fused shell), Lepidosaurs (lizards, snakes – venom, heat sensors), Crocodilians, Birds.
Birds: Feathers (keratin), hollow bones, one ovary, no teeth, endothermic; hard-shelled eggs; efficient flight metabolism.
Derived traits: mammary glands (milk), hair, fat insulation, kidneys, endothermy, large brain-to-body ratio, differentiated teeth, parental care.
Lineages:
Monotremes – Egg-laying (platypus, echidnas); lack nipples (milk on skin).
Marsupials – Pouched mammals (kangaroos, opossums, koalas); short gestation, embryonic young crawl to marsupium.
Eutherians (Placental mammals) – Long gestation, complex placenta (not detailed in transcript but implied).
Primates: Hands/feet with grasping, flat nails, large brain, forward eyes, fully opposable thumb (monkeys & apes); humans included.
Carnivores, Herbivores, Omnivores, Detritivores (e.g., earthworms), Scavengers, Filter feeders (sponges).
Meristem: Active plant cell division; apical (primary) vs lateral (secondary).
Alternation of Generations: Plant life cycle alternates haploid gametophyte & diploid sporophyte.
Homospory vs Heterospory: One vs two spore types; seed plants are heterosporous.
Body Cavity (Coelom): Enables organ independence & hydrostatic function.
Percentage figures:
Invertebrates constitute 95\% of animal species.
Six angiosperm crops supply 80\% of human calories.
Crop rotation, organic fertilizer, nitrogen fixation – sustainability principles.
Ethylene gas – plant hormone; accelerates fruit ripening (“one bad apple”).
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