The fossil record (two related questions, T9)
The fossil record provides evidence of the extinction of species, the origin and diversification of new groups, and transitional fossils
The geologic record is divided into the: Archaean, Proterozoic, and the Phanerozoic Eons. (Precambrian = Hadean + Archaean + Proterozoic)
The limits of the fossil record is that it is biased and incomplete because fossilization requires burial in sediment → probability of fossilization and discovery of fossils is very low
Relative dating → Sedimentary strata reveal the relative ages of fossils (limits: gaps in sedimentary sequence and sediments can be moved by major land movement)
Absolute dating → Radiometric dating
Origin of life (three questions, T9)
Life likely originate on earth from a series of chemical reactions that ultimately led to the formation of the first living organisms
Conditions on early earth made the chemical processes needed for life possible
It is not yet possible to definitively describe the steps that led to the mergence of life
Hypothesized multi-step process for the formation of simple cells:
Abiotic synthesis of small organic molecules
Terrestrial origins (deep-sea hydrothermal vents, volcanoes, Miller-Urey)
Extraterrestrial/Panspermia (have been detected in comets and meteorites)
Polymerization of small organic molecules into organic polymers
Monomers to polymers
Formation of protocells
cell-like structures that form when organic molecules become enclosed within vesicles
Emergence of self-replicating molecules
RNA world hypothesis, suggests that RNA was the first self-replicated molecule bc its much simpler than DNA
GOE → Great Oxygenation Event (2.7-2.3 b.y.a.)
when O2 became accumulating in the atmospheres and oceans which dramatically altered Earth’s environment
Caused the extinction of many prokaryotic groups, marking the end of the Archaean Eon
Prokaryote metabolic diversity (T10)
Prokaryotic metabolic diversity:
Phototrophs
Chemotrophs
Autotrophs
Heterotrophs
O2 and Nitrogen Metabolism
Endosymbiotic theory (two questions, T11)
Endosymbiosis is a symbiosis between two species in which one organism lives inside another organism’s cells or tissue
The endosymbiont theory (endosymbiotic origin of eukaryotic cells)
Endosymbionts → proteobacteria (mitochondria) and cyanobacteria (chloroplasts/plastids)
Primary endosymbiosis (1°) → Prokaryotic cells are engulfed as endosymbionts by either prokaryotic or eukaryotic cells
Secondary endosymbiosis (2°) → Eukaryotic cells themselves become endosymbionts being taken up by other eukaryotic cells
Animal phylogeny (T19)
Key features of the phylogeny of extant animals:
All animals share a single common ancestor → an ancestral colonial flagellated protist
Ancestral colonial flagellated protist → single-celled organism that formed a colony and had a flagella (whip like structures used for movement)
Kindom Animalia constitutes clade Metazoa, (all multicellular animals)
Sponges are basal (primitive) animals in the phylogeny
Eumatazoa (“true animals”) is a clade of animals with true tissues (excludes sponges)
Most animal phyla belong to the clade Bilateria animals with bilateral symmetry (bilaterians) (mirror halves)
Bilaterians split into 3 major clades—all are invertebrates, except Chordata, which includes vertebrates (like us with backbones)
Origin of animals (T19-20)
The common ancestor of animals evolved 800-650 mya and was a flagellated eukaryote.
The earliest multicellular eukaryotes
Multicellular eukaryotes evolved in multiple lineages, giving rise to algae, plants, fungi, and animals.
Whole-body animal fossils date from the Ediacaran Period: the Ediacaran biota (635-540 mya) are diverse soft-bodied, mostly sessile marine organisms that disappeared by the Cambrian.
Cambrian explosion (535-525 mya): the origin of most major phyla of living bilaterian animals.
Hypotheses for Cambrian explosion: evolution of predation; incr. in oceanic O2; evolution of the Hox genes.
Animal development (two questions, T19)
Embryonic development sequence:
Diploid zygote undergoes a series of rapis mitotic cell divisions call cleavage, transforming the zygote into a solid ball of cells
Cleave transforms morula into hollow blastula (hallow ball)
Gastrulation → ‘C’ fold inwards, producing two layers of embryonic tissues (ectoderm/outer layer and endoderm/inner layer)
Embryonic development is conserved by Hox genes
crucial becasue they play a fundamental role in determining the body plan and segment identity during embryonic development
Hox genes proteins coordinates the development of various structures along the anterior-posterior axis, e.g. legs, antennae, and wings in fruit flies or the different types of vertebrae in humans
Animals exhibit two primary modes of development:
Direct development → animal after birth or emergence from an egg is a small version of its adult form, eg. humans
Indirect development → has ‘larvae’ stages morphological behavioural differences from the sexually mature adult stages, eg. butterfly
Animal groupings (T19-20)
Invertebrates: Typically exhibit indirect development, with stages such as larval forms that differ significantly from adult morphology.
Vertebrates: Generally display direct development, although some exceptions exist with species like amphibians, which undergo metamorphosis before reaching adulthood.
Body plan:
Symmetry, Asymetrical, Radial, Bilateral, Cephalization
Tissues → diploblastic (ecto and endo) are radial, triploblastic (ecto, endo, and meso (muscle)
Triploblastic body cavity: coelom, hemocoel, no body cavity)
coelom is a fluid-filled body cavity completely lined with mesoderm, found in animals like humans and worms to cushion and support organs.
hemocoel is a blood-filled cavity found in insects and arthropods, where blood flows freely around organs in an open circulatory system.
Characteristics of eukaryotic life cycles (6 questions, all relevant topics)
Alternation between haploid and diploid stages
Haploid = 1 set of chromosomes (n)
Diploid = 2 sets (2n)
Meiosis creates haploid cells from diploid ones
Important for sexual reproduction
Reduces chromosome number
Fertilization restores diploid state
Two haploid gametes fuse to form a zygote
Mitosis helps with growth and development
Happens in both haploid or diploid stages, depending on the species
Three types of cycles:
Animal-like → diploid-dominant (humans)
Fungi-like → haploid-dominant
Plant-like → alternation of generations (both stages are multicellular)
Porifera/Sponges (T21)
Sessile marine animals (don’t move, stay in one spot), lack true tissues, and have motile larval stages for reproduction
Cell types:
epidermal cells (outer)
porocytes (create pores for water to enter)
choanocytes: beat flagella to create a water current and trap food particles,
mesohyl (amoebocytes → distribute food and make skeletal elements and spicules/spongin for structure)
Water enters via ostia (pores), flows into the spongocoel (central cavity), and exits through the osculum, choanocytes trap and engulf food particles by phagocytosis
Digested food either stays or gets passed to amoebocytes for distribution
Reproduction:
Hermaphrodites (both male and female organs)
Internal fertilization: eggs in mesohyl, fertilized by sperm
Free-swimming ciliated larvae disperse to form new sponges
Embryonic development is variable
Cnidaria (T21)
Cnidarians have two tissue layers: epidermis (outer) and gastrodermis (inner).
Their body is radially symmetrical with a central cavity that serves as both mouth and anus.
Tentacles surround the cavity, containing stinging cells (cnidocytes) with nematocysts to sting prey or defend.
Two body forms:
Polyp: Stationary (e.g., sea anemones).
Medusa: Bell-shaped and free-moving (e.g., jellyfish).
Cnidarians are predators, using tentacles to capture prey.
Life cycle and clades:
Medusozoans have a medusa stage.
Hydrozoans alternate between polyp and medusa forms.
Scyphozoans (jellyfish) and Cubozoans (box jellyfish) have the medusa form as the dominant stage.
Anthozoans (corals and sea anemones) exist only as polyps
Differentiating between invertebrates using characteristics (T21-25)
Lophotrochozoa:
Includes nearly half of all animal phyla (17 total)
Defined by molecular phylogeny; internal relationships remain unclear
Displays the widest range of body plans, from simple flatworms to complex octopuses
No single unifying characteristic across all members
Some develop a lophophore for feeding, others go through a trochophore larval stage, and a few have neither a lophophore nor a trochophore stage
Lophotrochozoa Phylums: Platyhelminthes, Syndermata, Ectoprocta, Brachiopoda, Throcozoa (Annelida, Mollusca)
Platylhelmintes (flatworms) → simple body plan, lack fluid filled body cavities, incomplete digestive tract (no anus), no circulatory or gas exchange system
Syndermata: Rotifers; tiny, pseudocoelomate, have a crown of cilia (corona) for feeding and movement, some are parasitic.
Ectoprocta: Bryozoans; colonial, sessile, have a lophophore (feeding structure), marine or freshwater, exoskeleton.
Brachiopoda: Lampshells; two hinged shells, lophophore for feeding, marine, not closely related to bivalves.
Annelida: Segmented worms; true coelom, closed circulatory system, segmented body, includes earthworms and leeches.
Mollusca: Soft-bodied with a hard shell (in most); have a mantle, muscular foot, and radula (except bivalves), includes snails, clams, and octopuses.
Ecdysozoa: group of animals that molt their exoskeleton (ecdysis); includes arthropods and nematodes
defined by molecular phylogeny
Cuticles → tough, non-living outer laters of organic material secreted outside the epidermis, protects against dissecation
Phylum Nematodes → roundworms
Arthropods: part of Ecdysozoa; segmented body, jointed limbs, exoskeleton made of chitin (e.g., insects, spiders, crustaceans), reduced coelom, open circulatory system
Deuterostomia: group where the anus forms before the mouth in development; includes chordates and echinoderms,
Chordates: part of Deuterostomia; animals with a notochord, dorsal nerve cord, and (usually) a backbone
Echinoderms: also Deuterostomes; marine animals with radial symmetry as adults (e.g., sea stars, sea urchins)
Parasitic invertebrates (T22)
Parasitic rhabitophorans live in or on other animals and are characterized by suckers/hooks for attachment to the host, tough outer coverings, and complex life cycles involving two or more hosts: intermediate host (asexual reproduction) and definitive host (sexual reproduction).
Important parasitic rhabditophorans incl. trematodes and tapeworms.
Trematodes parasitize a wide range of hosts, and most have complex life cycles with alternating sexual and asexual stages, e.g. blood flukes.