Mechanisms causing divergence
Selection
Non-random mating
Mutation
Genetic drift
Migration does not cause divergence
Biological species concept (BSC)
Reproductive isolation
Phylogenetic species concept (PSC)
Identify the smallest diagnosible monophyletic group
More useful and practical apparently
Both agree
Both agree on independent evolutionary units
Agree that evolutionary units lack gene flow
Disagree about the criteria used to determine when a lack of gene flow has happened
02/24/25
Speciation
1. Interrupt gene flow (migration)
Physical isolation (allopatric model)
Dispersal
getting past or going over a barrier
Vicariance (splitting)
A new barrier splits a current population
Glaciers, road construction
Sympatric model
Strong selection
Animals have a preference for extreme characteristics and ignore mates that don’t have this characteristic which can quickly drive a population to diverge
Reinforce speciation
Ignore
Behavioral isolation
Temporal
Habitat
Mechanical
Gametic (their gametes aren’t compatible)
2. Divergence
Mutation
Selection
Genetic drift
Non-random mating
Ecology
The study of species and species’ interactions with one another in their environment
Niche
Abiotic factors → combine to create a climate
Temperature
Water
Amount of sunlight
Wind
Soil
Biotic factors
02/26/25
Niche
Abiotic factors
Biotic factors
We can study niches in a more mathematical way by plotting them on a graph with the axes representing different variables
Competitive exclusion hypothesis
Two species cannot occupy the exact same niche in the exact same location, because then they would be in direct competition for the exact same set of resources
Partition of resources
Character displacement
Humans exterpated wolves from the United States and they eventually spread back into their niche after humans stopped interfering
02/28/25
Patters that drive community richness (diversity)
Island-equilibrium model
If the island is bigger, it should have more species on it
If two islands are the same size, the one that is closer to the mainland should have more species because animals can get there from the mainland more easily
As you remove islands you lose species because they can’t go somewhere else where their niche is already filled
Inter-specific species interactions
Predation (+/-)
One species eating another species
Causes co-evolution because each species is trying to outcompete the other
“Arms race”
Predator/prey: weapons → specialized structures, camouflage, strength, size, speed, intelligence, agility, endurance, social/solitary
Mutualism (+/+)
Both species are benefitting from the interactions
Oxpeckers, cleaner wray
parasitism/disease (+/-)
Parasitism: larger organisms that need boats (ticks, tapeworms)
Disease: smaller (parasites, bacteria)
Strengthens immune system, increases defensive mechanisms
Diseases and parasites evolve to avoid detection and avoid harming the host too much so as not to kill them
Commensalism (+/0)
One species benefits, the other is neutral
Not a very common species relationship
Herbivory (+/-)
Plants develop things like bigger size, spiny structures, and toxins
Animals develop the ability to eat them regardless
Trophic levels – trophic cascade!
Energy is lost as you go up the food chain
Dominant species is the most abundant species in the food web/chain
03/03/25
Dominant species
Very large biomass
Bottom-up control
Keystone species
Pivotal role in the community
Top-down control
Otters eat sea urchins so the kelp forests can grow
Population ecology
Intra-specific
∆N/∆T
dN/dT
Change in population in an interval of time
N = population size, T = time
dN/dT = r(N)
r = per capita rate of increase
d = per capita death rate
Ex: b = 10/100 = 0.1
Exponential growth curve, the population grows faster as time passes
03/10/25
Our relative abundance changes which impacts the resource availability for other species
~ 10,000 species of birds
~ 130 have gone extinct
~1% gone in 400 years
Maybe 100% gone in 40,000 years
Mass extinction – 60% of species in 1 million years