2.1 Atoms, Ions, and Molecules — Vocabulary Flashcards

2.1 a The chemical elements

  • Definition: An element is the simplest form of matter with unique chemical properties. Water is not an element; it is a compound made of elements hydrogen and oxygen.

  • Breakdown hierarchy:

    • Elements are composed of atoms.

    • Atoms contain protons, neutrons, and electrons; protons determine the atomic number (Z).

    • Protons and neutrons are in the nucleus; electrons form electron clouds around the nucleus.

  • Atomic number (Z): the number of protons in the nucleus. Examples:

    • Carbon: Z = 6

    • Oxygen: Z = 8

  • Periodic table arrangement: Elements are ordered by atomic number (Z).

  • Symbol notation: Elements are represented by one- or two-letter symbols. Examples:

    • C for carbon, Mg for magnesium, Na for sodium, Cl for chlorine.

    • Some symbols derive from Latin names, e.g., K for potassium (Kalium), Fe for iron (Ferrum).

  • Natural abundance in the human body:

    • There are 24 elements that play physiological roles.

    • Major elements (≈98.5% of body weight): O, C, H, N, Ca, P.

    • Next 0.8%: S, K, Na, Cl, Mg, Fe.

    • Trace elements: the remaining ≈0.7% overall.

  • Trace elements: essential in small amounts but can be toxic in excess (e.g., lead, mercury).

  • Minerals vs. elements:

    • Minerals are inorganic elements obtained from soil via plants; they contribute to structure and function.

    • Minerals constitute about 4% of body weight. Roughly 75% of these minerals are Ca, P, Mg, and K (often abbreviated CAMP).

  • Roles of minerals:

    • Bone and teeth: crystals of Ca, phosphate, Mg, fluoride, sulfate.

    • Proteins: sulfur-containing amino acids; phosphorus in nucleic acids, ATP, membranes.

    • Enzymes: minerals enable enzyme function; iodine in thyroid hormone; Fe in hemoglobin; Mn, Zn, Cu as enzyme cofactors.

  • Electrolytes: mineral salts that dissociate into ions in body fluids and are essential for nerve and muscle function; they influence water distribution and osmotic balance.

  • Practical/clinical notes:

    • Electrolyte balance is critical in patient care; imbalances can cause cramps, bone brittleness, coma, or cardiac arrest.

    • Heavy metal poisoning (e.g., Pb, Hg) disrupts physiology.

2.1 b Atomic structure

  • Historical context:

    • Democritus (5th c. BCE) proposed atoms as indivisible particles.

    • Dalton (early 1800s) developed an experimental atomic theory.

    • Bohr (1913) proposed a planetary model; later quantum mechanics showed electrons occupy probabilistic regions rather than fixed orbits.

  • Nucleus:

    • Center of the atom contains protons (p+) and neutrons (n0).

    • Protons carry a positive charge; neutrons are neutral.

    • Each proton/neutron ~ 1 atomic mass unit (amu).

    • Atomic mass roughly equals the total number of protons and neutrons (A ≈ Z + N).

  • Electrons:

    • Electrons (e−) have a negative charge and very small mass; mass of electrons is negligible compared to protons/neutrons for most purposes (≈ 1/1836 of a proton).

    • Electrons occupy electron shells around the nucleus; shells have energy levels and hold a limited number of electrons.

    • The number of electrons equals the number of protons for electrical neutrality: Z electrons for a neutral atom.

  • Most physiologically relevant elements involve up to four electron shells (despite that some atoms can have up to seven).

  • Valence electrons:

    • Outer-shell electrons (valence electrons) determine chemical bonding properties of an atom.

  • Visual note: Bohr’s model is a useful visualization, but not a literal depiction of atomic structure in quantum terms.

2.1 c Isotopes and radioactivity

  • Isotopes: variants of a given element with the same number of protons (Z) but different numbers of neutrons (N); thus different atomic masses.

    • Hydrogen isotopes: protium 1H (1 p+), deuterium 2H (1 p+, 1 n), tritium 3H (1 p+, 2 n).

    • Carbon isotopes: 12C (6 p+, 6 n), 13C (7 n), 14C (8 n).

  • Chemical behavior: all isotopes of a given element generally behave the same chemically; differences are physical (e.g., mass, stability).

  • Example: Deuterium (2H) reacts with oxygen to form water just like ordinary hydrogen (1H).

  • Atomic weight vs. isotopic composition:

    • Atomic weight is the average mass of all isotopes weighted by their natural abundance (e.g., carbon’s atomic weight ≈ 12.011 amu due to heavier isotopes 13C, 14C).

  • Radioactivity:

    • Some isotopes are unstable and decay to more stable isotopes; this process emits high-energy radiation (ionizing radiation).

    • Common decay modes: alpha (α) particles, beta (β) particles, gamma (γ) rays.

    • An alpha particle is a helium nucleus (2 protons, 2 neutrons).

    • A beta particle is an electron.

    • Gamma rays are high-energy photons.

  • Half-life concepts:

    • Physical half-life: time required for 50% of a radioisotope to decay.

    • Example physical half-lives: 90Sr ≈ 28 years; 40K ≈ 1.3 × 10^9 years.

    • Biological half-life: time required for half of a radioisotope to disappear from the body (through decay plus excretion).

    • Example: Cs-137 has physical half-life ≈ 30 years but a biological half-life ≈ 17 days due to rapid excretion.

  • Health effects:

    • Ionizing radiation can eject electrons, create free radicals, and damage biomolecules (DNA), leading to mutagenesis and carcinogenesis.

    • High doses can be fatal; lower doses pose risks over time.

  • Measurements and units (brief):

    • Dose concepts are measured in sieverts (Sv) to reflect biological effect, accounting for radiation type and tissue sensitivity.

    • In occupational settings, limits are specified (e.g., around 50 mSv/year).

2.1 d Ions, electrolytes, and free radicals

  • Ions:

    • Charged atoms or molecules; cations are positively charged; anions are negatively charged.

    • Result from electron transfer during ionization or from chemical bonding in solution.

    • Example ion pair: Na⁺ (cation) and Cl⁻ (anion).

    • Some elements have multiple ionic states (e.g., iron: Fe²⁺ = ferrous, Fe³⁺ = ferric).

    • The charge on an ion is called its valence.

  • Electrolytes:

    • Substances that dissociate into ions in water; essential for chemical reactivity, osmotic balance, and electrical phenomena in nerves and muscles.

    • Common electrolytes include ions such as Na⁺, K⁺, Ca²⁺, Mg²⁺, Cl⁻, HCO₃⁻, and others.

    • Electrolytes enable many physiological processes, including nerve conduction and muscle contraction.

  • Free radicals:

    • Unstable, highly reactive chemical species with odd (unpaired) electrons, e.g., superoxide anion $ ext{O}_2^{ullet-}$.

    • Sources: normal metabolic reactions (mitochondrial oxidation), immune defense, and exposure to radiation or certain chemicals.

    • Dangers: react with fats, proteins, and DNA to form damaging radicals; implicated in aging and diseases such as cancer and heart disease.

  • Antioxidants and protection:

    • The body has natural antioxidant defenses, including the enzyme superoxide dismutase (SOD), which converts superoxide to oxygen and hydrogen peroxide.

    • Dietary antioxidants include selenium, vitamin E (α-tocopherol), vitamin C (ascorbic acid), and carotenoids (e.g., β-carotene).

2.1 e Molecules and chemical bonds

  • Molecules vs. compounds:

    • Molecule: two or more atoms bound together.

    • Compound: a molecule composed of at least two different elements.

    • Example: $ ext{O}2$ (molecule) vs. $ ext{CO}2$ (molecule and compound).

  • Isomers:

    • Molecules with the same molecular formula but different structures (e.g., ethanol vs. ethyl ether; both $ ext{C}2 ext{H}6 ext{O}$).

  • Molecular weight (MW): sum of atomic weights of all atoms in the molecule.

    • Glucose: $ ext{C}6 ext{H}{12} ext{O}_6$ → $MW
      oughly = 6(12) + 12(1) + 6(16) = 180$ amu.

  • Bonds of greatest physiological interest:

    • Ionic bonds, covalent bonds, hydrogen bonds, and van der Waals forces.

  • Ionic bonds:

    • Attraction between cations and anions (e.g., Na⁺ and Cl⁻) forming salts like NaCl.

    • Ionic compounds tend to dissociate in water because ions are more attracted to water than to each other.

    • Question to consider: Are ionic bonds common in human body fluids? (Typically, salts exist as dissociated ions in body fluids rather than as intact ionic bonds in solution.)

  • Covalent bonds:

    • Single covalent bonds: sharing one pair of electrons (e.g., H–H).

    • Double covalent bonds: sharing two pairs of electrons (e.g., O=C=O in CO₂).

    • Nonpolar covalent bonds: electrons spent evenly around both nuclei.

    • Polar covalent bonds: electrons spend more time near one nucleus (e.g., O–H in water) creating partial charges.

  • Hydrogen bonds:

    • A weak attraction between a slightly positive hydrogen in one molecule and a slightly negative atom (usually O or N) in another or in the same molecule.

    • Essential for water's properties and for maintaining 3D structures of large biomolecules (proteins, DNA).

    • Represented by dotted lines in diagrams.

  • Van der Waals forces:

    • Weak, brief attractions between neutral atoms due to transient dipoles caused by electron density fluctuations.

    • Individually weak but can be significant collectively (e.g., protein folding, membrane interactions, gecko adhesion).

2.2 water and mixtures

2.2 a Water

  • Water structure and polarity:

    • Water is a polar molecule with a V-shaped geometry; bond angle ~ $105^ ext{o}$.

    • Oxygen bears a partial negative charge ($
      abla^- ext{ delta minus}$) and hydrogens bear partial positive charges ($
      abla^+ ext{ delta plus}$).

  • Key properties arising from polarity and hydrogen bonding:

    • Solvency: water dissolves many substances; water is the universal solvent.

    • Hydrophilic vs. hydrophobic:

    • Hydrophilic: substances that readily dissolve in water (e.g., NaCl).

    • Hydrophobic: substances that do not dissolve well in water (e.g., fats).

    • Cohesion: water molecules stick to each other via hydrogen bonds; explains surface tension and puddle formation.

    • Adhesion: water’s tendency to cling to surfaces, aiding lubrication and movement of certain tissues.

    • Chemical reactivity: water participates in reactions (hydrolysis and dehydration synthesis) and can ionize into $ ext{H}^+$ and $ ext{OH}^-$.

    • Thermal stability: high heat capacity; many reactions and temperature regulation rely on water’s ability to absorb heat without large temperature changes. When water vaporizes, it carries a lot of heat away, cooling the body (e.g., perspiration).

  • Hydration and solvation:

    • Hydration spheres form around ions in solution: e.g., Na⁺ ions attract water with the oxygen end; Cl⁻ ions attract water with the hydrogen ends.

  • Roles in physiology:

    • Transport medium for dissolved substances; solvent for metabolic reactions.

2.2 b Solutions, colloids, and suspensions

  • Mixtures vs. compounds:

    • Mixture: substances physically blended but not chemically combined; each retains properties.

    • Compound: elements chemically reacting to form a substance with new properties (e.g., NaCl).

  • Types of mixtures in the body:

    • Solution: particles < 1 nm; transparent; solute passes through most membranes; particles do not settle; examples include saline in plasma, glucose in blood.

    • Colloid: particles 1–100 nm; cloudy; do not settle; cannot pass through most membranes; examples include proteins in plasma.

    • Suspension: particles > 100 nm; cloudy or opaque; particles settle; examples include blood cells in plasma.

    • Emulsion: a suspension of one liquid in another (e.g., fats in breast milk, oil in vinegar). Blood can be described as a combination: a solution (NaCl), a colloid (protein), and a suspension (cells).

  • Table 2.4 (types of mixtures) provides representative examples for the body.

2.2 c Acids, bases, and pH

  • Definitions:

    • Arriving at acid/base concepts via proton transfer:

    • Acid: proton donor (releases H⁺ in water).

    • Base: proton acceptor (accepts H⁺).

    • Ammonia (NH₃) is a base because it accepts a proton to become NH₄⁺.

  • pH concept:

    • pH is the negative logarithm of the hydrogen ion concentration: ext{pH} = - ext{log}_{10}ig[H^+ig]

    • Pure water:

    • At equilibrium, $[H^+][OH^-] = 10^{-14}$ (at 25°C).

    • In pure water, each is $10^{-7} ext{ M}$, giving pH = 7 (neutral).

    • pH scale: 0–14; below 7 is acidic, above 7 is basic (alkaline).

    • The lower the pH, the higher the hydrogen ion concentration; the scale is logarithmic, so a change of 1 pH unit corresponds to a tenfold change in $[H^+]$.

  • Physiological relevance and buffers:

    • Blood normally maintained around pH 7.35–7.45; deviations disrupt physiology.

    • Buffers resist pH changes; discussed in detail in later chapters.

  • pH and drug action (clinical insight):

    • The pH of body fluids affects drug ionization and membrane permeability (e.g., aspirin in the stomach is uncharged and can cross membranes; in the bloodstream it ionizes and may get trapped).

    • Concept of ion trapping (pH partitioning) can be used to enhance excretion of poisons by manipulating urine pH.

2.2 d The other measures of concentration

  • Concentration units:

    • Weight per volume: grams or milligrams per liter (e.g., serum cholesterol 200 mg/dL).

    • Percentage: w/v (weight/volume) or v/v (volume/volume).

    • Molarity (M): number of moles of solute per liter of solution; reflects molecule number rather than mass.

  • Millimolar and milliequivalents:

    • Body fluids are often quantified in millimolar (mM) concentrations to reflect actual molar amounts.

    • Milliequivalents per liter (mEq/L) account for electrical charge of ions, important for nerve and muscle physiology.

  • Practical importance: units of concentration influence interpretation of electrolyte balance and IV fluid composition.

2.3 energy chemical reactions

2.3 a Energy and work

  • Definitions:

    • Energy: capacity to do work.

    • Work: movement of a physical object or, in physiology, movement of molecules/ions or mechanical processes (e.g., pumping blood, muscle contraction).

  • Energy forms:

    • Chemical energy: stored in chemical bonds; released during reactions to power physiological work.

    • Heat: kinetic energy of molecular motion; temperature measures the rate of motion.

    • Electromagnetic energy: moving quanta (photons), e.g., light.

    • Electrical energy: moving charges; potential when charges are separated (e.g., membrane potentials); kinetic when ions move to produce currents.

  • Free energy (G): the energy available to do useful work in a system.

  • In physiology, the primary free energy source is the chemical energy stored in bonds of organic molecules.

2.3 b Classes of chemical reactions

  • Chemical reactions involve bond formation or breaking; symbolized by chemical equations with reactants → products.

  • Examples:

    • Oxidation of ethanol by oxygen to form acetic acid and water (wine turning into vinegar).

  • Reaction types:

    • Decomposition: large molecules break into smaller pieces (e.g., starch → glucose).

    • Synthesis: smaller molecules combine into larger ones (e.g., amino acids into proteins).

    • Exchange: atoms or groups are exchanged between molecules (e.g., HCl neutralization in digestion).

  • Reversibility:

    • Many reactions are reversible and proceed in either direction depending on conditions.

    • Law of mass action governs direction: reactions proceed from higher to lower reactant/product concentrations.

    • Biochemical example: CO₂ + H₂O ⇌ H₂CO₃ ⇌ HCO₃⁻ + H⁺; equilibrium can shift with changes in CO₂, bicarbonate, or H⁺.

  • Equilibrium concept:

    • Under normal conditions, the carbonic acid/bicarbonate system maintains a physiological balance (e.g., CO₂ transport and blood pH).

2.3 c Reaction rates

  • Factors affecting reaction rates:

    • Concentration: higher reactant concentration → more frequent collisions → faster rate.

    • Temperature: higher temperature → faster molecular motion → more frequent/energetic collisions.

    • Catalysts: substances that increase reaction rate by providing a favorable orientation or lowering activation energy; are not consumed in the reaction.

  • Biological catalysts:

    • Enzymes are the most important biological catalysts; they speed up reactions under physiological conditions.

2.3 d Metabolism, oxidation, and reduction

  • Metabolism: all chemical reactions in the body.

    • Subdivisions: catabolism (energy-releasing) and anabolism (energy-storing).

    • Exergonic reactions release energy; endergonic reactions require energy input.

  • Oxidation-reduction (redox) reactions:

    • Oxidation: loss of electrons (molecule becomes more positive/less reduced); the oxidizing agent is the electron acceptor.

    • Reduction: gain of electrons (molecule becomes more reduced); the reducing agent is the electron donor.

    • In many cases electrons are transferred as hydrogen atoms; the presence of oxygen as electron acceptor is a common reason for using the term oxidation.

  • Redox couple notation:

    • A− → A is a reducing agent; B is oxidized; E− represents the transferred electrons.

  • Links to energy transfer:

    • Energy from exergonic oxidation reactions powers endergonic synthetic reactions in metabolism.

2.4 organic compounds

2.4 a Monomers and polymers

  • Organic chemistry basics:

    • Carbon is unique due to four valence electrons, allowing diverse bonding and the formation of long chains, branched structures, and rings.

    • Carbon forms covalent bonds with H, O, N, S; rich functional groups determine properties.

  • Polymers and monomers:

    • Macromolecules consist of repeating subunits called monomers.

    • Examples: starch (polysaccharide of glucose monomers) and proteins (polypeptides composed of amino acids).

  • Polymerization and hydrolysis:

    • Dehydration synthesis (condensation): monomers join, releasing water, forming covalent bonds and a polymer chain.

    • Hydrolysis: a water molecule breaks covalent bonds, yielding monomers; enzymes catalyze the process.

2.4 b Carbohydrates

  • General formula: carbohydrates have the form $CH2O$ repeating units; commonly written as $(CH2O)_n$.

  • Important monosaccharides: glucose, galactose, fructose; all C₆H₁₂O₆ isomers.

  • Other sugars:

    • Ribose (RNA) and deoxyribose (DNA).

  • Disaccharides: two monosaccharides linked, e.g., sucrose (glucose + fructose), lactose (glucose + galactose), maltose (glucose + glucose).

  • Polysaccharides: long chains of monosaccharides; 10–20 units can be oligosaccharides; 50+ units typically polysaccharides.

    • Glycogen: animal storage polysaccharide; liver, muscle, brain store glycogen; highly branched.

    • Starch: plant storage polysaccharide; digestible in humans.

    • Cellulose: structural polysaccharide in plants; not digestible by humans but provides dietary fiber.

  • Conjugated carbohydrates:

    • Glycolipids and glycoproteins: carbohydrates covalently bound to lipids or proteins; roles in cell membranes and mucus (protection, infection resistance).

  • Glycoconjugates:

    • Proteoglycans: protein–carbohydrate complexes important for gels that hold tissues together and lubricate joints; components in connective tissues and eye.

  • Functions of carbohydrates:

    • Major energy source; glucose oxidation generates ATP.

    • Some carbohydrates serve as cell surface markers and structural components.

2.4 c Lipids

  • General features:

    • Lipids are hydrophobic, with high hydrogen-to-oxygen ratio; mainly carbon, hydrogen, and oxygen.

    • Energy density is high (more calories per gram than carbohydrates).

  • Major lipid types in humans:

    • Fatty acids: carboxyl group at one end and methyl group at the other; saturated vs. unsaturated; polyunsaturated fats have multiple C=C bonds.

    • Triglycerides (neutral fats): glycerol backbone with three fatty acids; energy storage form; fats (solid at room temperature) vs oils (liquid at room temperature).

    • Phospholipids: glycerol + two fatty acids + a phosphate-containing head; amphipathic; major components of cell membranes; lecithin (phosphatidylcholine) as a common phospholipid.

    • Eicosanoids: 20-carbon signaling molecules derived from arachidonic acid; include prostaglandins involved in inflammation, clotting, hormone action, uterine contractions, and more.

    • Steroids: lipids with four fused rings; cholesterol is the main steroid, precursor to other steroids (cortisol, estrogens, testosterone) and bile acids; cholesterol is essential for membranes and nerve function; about 15% of cholesterol is from the diet; 85% is synthesized endogenously, mainly by the liver.

  • Cholesterol discussion:

    • LDL (low-density lipoprotein) is often labeled as “bad” cholesterol due to association with atherosclerosis; HDL (high-density lipoprotein) is considered “good” as it helps remove cholesterol from the bloodstream.

    • Trans fats: partially hydrogenated fats with trans double bonds; denser packing leads to higher risk of cardiovascular disease; FDA banned artificial trans fats in 2015 due to health concerns.

  • Practical notes:

    • Main roles of fats: energy storage, insulation, cushioning of organs; phospholipids form cell membranes; prostaglandins regulate various physiological processes.

2.4 d Proteins

  • Proteins overview:

    • Proteins are the most versatile biological macromolecules; they perform structural, catalytic, signaling, transport, and protective roles.

    • A protein is a polymer of amino acids.

  • Amino acids:

    • 20 standard amino acids share the same backbone: an α-carbon attached to an amino group (–NH₂), a carboxyl group (–COOH), a hydrogen, and a variable R-group.

    • R-groups determine chemical properties (hydrophilic vs. hydrophobic) and function.

    • The amino acid sequence is indicated by the protein’s primary structure.

  • Peptides and polypeptides:

    • A peptide is a molecule with two or more amino acids joined by peptide bonds (dehydration synthesis).

    • Dipeptide (2 aa), tripeptide (3 aa); oligopeptides (<≈15–20 aa); polypeptides (>≈50 aa).

  • Protein structure levels:

    • Primary structure: amino acid sequence encoded by genes.

    • Secondary structure: regular folding patterns stabilized by hydrogen bonds, mainly alpha helices and beta sheets.

    • Tertiary structure: overall 3D folding driven by hydrophobic/hydrophilic interactions, hydrogen bonds, ionic interactions, and van der Waals forces.

    • Quaternary structure: assembly of multiple polypeptide chains (e.g., hemoglobin with four subunits).

  • Factors stabilizing structure:

    • Hydrogen bonds, ionic bonds, hydrophobic interactions, and van der Waals forces.

    • Disulfide bridges (covalent) between cysteine residues can stabilize tertiary and quaternary structures.

  • Protein function categories:

    • Structure: keratin, collagen.

    • Communication: hormones, receptors, signaling ligands.

    • Membrane transport: channels and carriers (e.g., Na⁺/K⁺-ATPase).

    • Catalysis: enzymes; most metabolic pathways are enzyme-catalyzed.

    • Recognition and protection: antibodies and other immune proteins.

    • Movement: motor proteins (e.g., myosin).

    • Adhesion: proteins that bind cells to each other.

  • Enzymes (2.4 f):

    • Enzymes are biological catalysts; most are proteins; some ribozymes (RNA enzymes) exist in some contexts.

    • Names: traditionally named after substrate (e.g., amylase acts on starch); modern naming uses suffix -ase and may include the substrate or class (e.g., carbonic anhydrase).

    • Active site: the region where substrates bind; enzyme–substrate complex forms via induced fit, where the enzyme changes shape to accommodate the substrate.

    • Enzyme specificity: each enzyme acts on a particular substrate; some enzymes catalyze two substrates at two active sites.

    • Denaturation: extreme conditions (temperature, pH) disrupt hydrogen bonds and other weak forces, altering enzyme shape and function.

    • Cofactors and coenzymes:

    • Cofactors: inorganic ions (Fe, Cu, Zn, Mg, Ca, etc.) that assist enzyme activity.

    • Coenzymes: organic molecules (often derived from vitamins) that transfer electrons or functional groups (e.g., NAD⁺, FAD).

  • Metabolic regulation and isoenzymes:

    • Isoenzymes (esoenzymes) are enzyme variants with the same function but different structures or regulatory properties; they help in diagnosing diseases (e.g., CK-MM for skeletal muscle, CK-MB for heart).

  • Enzymes in metabolism example:

    • Carbonic anhydrase hydrolyzes carbonic acid to CO₂ and water at very high rates (e.g., up to ~3.6×10^7 molecules per minute for a single enzyme).

  • Enzyme mechanisms and examples:

    • Sucrose breakdown by the enzyme sucrase (invertase) is an example of substrate binding to an active site and cleavage in the presence of water (hydrolysis).

2.4 g ATP, other nucleotides, and nucleic acids

  • Nucleotides:

    • Basic components: a nitrogenous base, a monosaccharide (ribose or deoxyribose), and one or more phosphate groups.

    • ATP (adenosine triphosphate) is a nucleotide with adenine base, ribose sugar, and three phosphates; a key energy-transfer molecule.

  • ATP in energy transfer:

    • ATP briefly stores energy from exergonic reactions and releases it for work (e.g., polymerization, muscle contraction, ion transport).

    • The high-energy bonds linking the phosphate groups are typically represented with a wavy line in structural formulas; hydrolysis releases energy.

    • Hydrolysis of the terminal phosphate bond yields ADP and inorganic phosphate (Pi):

    • Reaction: ext{ATP}
      ightarrow ext{ADP} + ext{P}_i + ext{energy} \ ext{Energy release} \ ext{~7.3 kcal/mol}

    • The energy release is used for cellular work; not all energy released is used as work—much is released as heat.

  • phosphorylation:

    • The free phosphate released can be transferred to enzymes or other molecules to activate them; this process is called phosphorylation and is carried out by kinases.

  • ATP turnover and energy balance:

    • The body consistently replenishes ATP; under moderate activity, a full day’s supply would weigh roughly 45 kg (about 99 lb).

    • The continuous need for ATP is why mitochondria are considered the cell’s powerhouses, where aerobic respiration occurs to regenerate ATP.

  • Other nucleotides and signaling:

    • GTP (guanosine triphosphate) can donate phosphate groups to ADP to regenerate ATP in some contexts.

    • cAMP (cyclic adenosine monophosphate) acts as a second messenger in many signaling pathways.

  • Nucleic acids:

    • DNA (deoxyribonucleic acid): polymers of nucleotides; carries genetic information; typically 100,000,000 to 1,000,000,000 nucleotides long.

    • RNA (ribonucleic acid): ribonucleotides; carries out coding and synthesis tasks; three major forms (mRNA, tRNA, rRNA) with lengths ranging from ~70 to ~10,000 nucleotides in various contexts.

  • Overview of nucleic acids in biology:

    • DNA stores genetic information and provides instructions for protein synthesis; transmission during cell division and heredity.

    • RNA translates genetic information into proteins and catalyzes synthesis in conjunction with ribosomes.

2.4 h Nucleic acids (continuation)

  • The genetic code and protein synthesis are detailed in Chapter 4, but the essential idea is:

    • DNA sequences encode amino acid sequences in proteins via transcription to mRNA and translation on ribosomes.

    • A, G, C, T (DNA) or U (RNA) nucleotides form the language of genetics.

2.4 f Enzymes in metabolism (summary)

  • Enzymes are biological catalysts that accelerate metabolic reactions under physiological conditions.

  • Cofactors and coenzymes expand the range of reactions that enzymes can catalyze and regulate their activity.

  • Enzyme activity is influenced by temperature, pH, and the presence of activators or inhibitors.

  • The concept of activation energy: even exergonic reactions require a minimum energy input to overcome the energy barrier; enzymes lower this barrier, allowing reactions to proceed rapidly at body temperature.

2.4 g Clinical insights and broader context

  • Trans fats and cardiovascular health:

    • Trans fats (partially hydrogenated oils) package fats in a way that resists enzymatic breakdown, remain in circulation longer, and tend to deposit in arteries.

    • Large studies linked higher trans fat intake to increased risk of coronary artery disease (CAD).

    • FDA banned artificial trans fats in processed foods due to health concerns.

  • Good vs. bad cholesterol:

    • LDL (low-density lipoprotein) is often labeled as 'bad' cholesterol; HDL (high-density lipoprotein) as 'good' cholesterol.

    • LDL contributes to atherosclerosis when elevated; HDL helps remove cholesterol from circulation.

  • Steroid use and health risks:

    • Anabolic/androgenic steroids (synthetic testosterone derivatives) can promote muscle growth but carry significant health risks: liver damage, cardiovascular disease, immune suppression, hormonal imbalances, and psychiatric effects.

    • Societal and ethical concerns accompany steroid use in sports.

  • Medical and physiological implications:

    • The body’s lipid profile, cholesterol balance, and steroid metabolism affect health; understanding lipids and proteins helps explain disease processes and treatments.

Quick conceptual recap (glossary-style)

  • Atom vs. element vs. isotope: atoms form elements; isotopes differ in neutron count but share chemical properties; radioisotopes are unstable and radioactive.

  • Ions and electrolytes: ions carry charges; electrolytes dissociate in water and enable electrical signaling and fluid balance.

  • Water’s role in life: solvent, thermal buffer, participant in reactions, and facilitator of transport.

  • Macromolecules: carbohydrates, lipids, proteins, nucleic acids; monomers → polymers via dehydration synthesis; water-driven hydrolysis = digestion.

  • Bonding types: ionic, covalent (polar and nonpolar), hydrogen bonds, van der Waals forces; all contribute to biomolecular structure and function.

  • Energy and metabolism: chemical energy stored in bonds; ATP as the primary energy currency; catabolism vs. anabolism; oxidation-reduction in energy transfer.

  • Enzymes and regulation: specificity and induced fit; cofactors and coenzymes; isoenzymes used in diagnostics.

  • pH and buffers: acid-base balance in the body; pH affects molecule ionization and consequently drug action and physiology.

  • Clinical relevance: trans fats, cholesterol transport, isotope usage, radiological safety, and enzyme-based diagnostics.

  • Note: The content above consolidates the major and supporting points from sections 2.1–2.4, including the examples, definitions, and implied clinical implications discussed in the transcript.