Evolution
Chapter 22: Descent with Modification
Relative dating is a method of comparing the ages of two fossils by comparing their placement in rock layers
Artificial selection is the selective breeding of plants and animals to produce animals to produce desirable traits in offspring
Inheritance of Acquired Characteristics: modifications can be passed on to offspring
Natural Selection: the process by which organisms with variation most suited to their environment to survive and leave more offspring
Fitness: How well an organism can survive and reproduce in its environment.
Evidence for Evolution:
Direct Observation
Ex. Insect populations become resistant to pesticides Antibiotic-resistant bacteria
Fossil Record
Homology: characteristics in related species can have underlying similarities even though functions may differ
Ex. Homologous structures: similar anatomy from common ancestors
Embryonic homologies: similar early development
Vestigial Organs: Structures with little to no use
Molecular homologies: similar DNA and amino acid
Biogeography: Geographic distribution of a species
Ex. Continental drift and Pangaea explain similarities on different continents.
Convergent evolution: Distantly related species can resemble one another Similar problem, similar solutions
Ex. Torpedo shape of sharks, penguins, and dolphins
Divergent Evolution: Where you diverge/move away from common ancestors
Ex. Humans diverging from the monkeys
Chapter 23: Evolution of Populations
Microevolution: change in allele frequencies of a population over generations
Evolution is based on genetic variation
Point mutations: changes in one base (eg. sickle cell)
Chromosomal mutations: delete, duplicate, disrupt, rearrange
Sexual recombination: contributes to most of the genetic variation in a population (Crossing over, independent assortment, random fertilization)
Population genetics: study of how populations change genetically over time
Population: a group of individuals that live in the same area and interbreed, producing fertile offspring
Gene pool: all of the alleles for all genes in all the members of a population
Fixed allele: all members of a population only have 1 allele for a particular trait
Hardy-Weinberg Principle: the allele and genotype frequencies of a population will remain constant from generation to generation
Equilibrium + allele and genotype frequencies remain constant
CONDITIONS FOR HARDY-WEINBERG EQUILIBRIUM
No mutations
Random mating
No natural selection
Large population size
No gene flow
If at least one of these conditions is NOT met, then the population is EVOLVING
Allele frequencies: a gene with 2 alleles: p, q
p = frequency of dominant allele (A)
q = frequency of recessive allele (a)
p + q = 1
USE DECIMALS
Genotype frequencies: 3 genotypes (AA, Aa, aa)
p2 + 2pq + q2 = 1
p2 = AA (Homozygous dominant)
2pq = Aa (Heterozygous)
q2 = aa (Homozygous recessive)
If you are given the genotypes, calculate p and q by adding up the total # of A and a alleles
If you know phenotypes, then use “aa” to find q2, and then q. (p = q-1)
Use p2 + 2pq + q2 to find genotype frequencies
The key to solving H-W problems lies with the homozygous recessive individuals
Minor Causes:
Mutations: Rare, very small changes in allele frequencies
Nonrandom Mating: Affect genotypes, but not allele frequencies
Major Causes: Natural Selection, genetic drift, gene flow
Founder Effect: A few individuals isolated from larger populations
Bottleneck Effect: Sudden change in the environment drastically reduces population size
3 types of Natural Selection:
Directional Selection
Disruptive (diversifying) Selection
Stabilizing Selection
Sexual Selection: Form of natural selection - certain individuals more likely to obtain mates
Sexual Dimorphism: Difference between 2 sexes - size, color, ornamentation, behavior
Chapter 24: Speciation
Speciation: Origin of species
Microevolution: Changes within a single gene pool
Macroevolution: Evolutionary changes that cause one species to turn into 2 different species
Reproductive isolation: barriers that prevent members of 2 species from producing viable, fertile offspring
Prezygotic Barriers:
Impede Mating/Fertilization
Ex. Habitat Isolation, Temporal Isolation, Behavioral Isolation, Mechanical Isolation, Gametic Isolation
Postzygotic Barriers:
Prevent hybrid zygote from developing into viable adult
Ex. Reduced Hybrid Viability, Reduced Hybrid Fertility, Hybrid Breakdown
Allopatric Speciation: A population forms a new species while geographically isolated from its parent population
Sympatric Speciation: A subset of a population forms a new species without geographic separation
***Polyploidy means instant speciation (for plants)***
Adaptive Radiation: Many new species arise from a common ancestor
Occurs when:
A few organisms make a way to new, distant areas (allopatric speciation)
Environmental Change -> Extinctions -> New Niches for Survivors
Ex. Hawaiian Archipelago
Hybrid Zones:
Incomplete Reproductive Barriers
Possible Outcomes: reinforcement, fusion, stability
Tempo of Evolution:
Gradualism
Common Ancestor
Slow, Constant Change
Punctuated Equilibrium:
Eldridge & Gould
A long period of stasis punctuated by short bursts of significant change
Chapter 26: Cladistics/Phylogeny
Tools used to determine evolutionary relationships:
Fossils
Morphology (homologous structures)
Molecular Evidence (DNA, amino acids)(DNA does not lie)
Phylogenetic Tree: branching diagram that shows the evolutionary history of a group of organisms
***Nodes show where a common ancestor is between organisms***
Cladogram: Diagram that depicts patterns of shared characteristics among taxa
Clade: group of species that includes ancestral species + all descendants
Shared derived characteristics are used to construct cladograms
***Branch lengths can indicate time***
3 Domains: Bateria, Archea, Eukarea
***Nodes can be rotated***
Chapter 22: Descent with Modification
Relative dating is a method of comparing the ages of two fossils by comparing their placement in rock layers
Artificial selection is the selective breeding of plants and animals to produce animals to produce desirable traits in offspring
Inheritance of Acquired Characteristics: modifications can be passed on to offspring
Natural Selection: the process by which organisms with variation most suited to their environment to survive and leave more offspring
Fitness: How well an organism can survive and reproduce in its environment.
Evidence for Evolution:
Direct Observation
Ex. Insect populations become resistant to pesticides Antibiotic-resistant bacteria
Fossil Record
Homology: characteristics in related species can have underlying similarities even though functions may differ
Ex. Homologous structures: similar anatomy from common ancestors
Embryonic homologies: similar early development
Vestigial Organs: Structures with little to no use
Molecular homologies: similar DNA and amino acid
Biogeography: Geographic distribution of a species
Ex. Continental drift and Pangaea explain similarities on different continents.
Convergent evolution: Distantly related species can resemble one another Similar problem, similar solutions
Ex. Torpedo shape of sharks, penguins, and dolphins
Divergent Evolution: Where you diverge/move away from common ancestors
Ex. Humans diverging from the monkeys
Chapter 23: Evolution of Populations
Microevolution: change in allele frequencies of a population over generations
Evolution is based on genetic variation
Point mutations: changes in one base (eg. sickle cell)
Chromosomal mutations: delete, duplicate, disrupt, rearrange
Sexual recombination: contributes to most of the genetic variation in a population (Crossing over, independent assortment, random fertilization)
Population genetics: study of how populations change genetically over time
Population: a group of individuals that live in the same area and interbreed, producing fertile offspring
Gene pool: all of the alleles for all genes in all the members of a population
Fixed allele: all members of a population only have 1 allele for a particular trait
Hardy-Weinberg Principle: the allele and genotype frequencies of a population will remain constant from generation to generation
Equilibrium + allele and genotype frequencies remain constant
CONDITIONS FOR HARDY-WEINBERG EQUILIBRIUM
No mutations
Random mating
No natural selection
Large population size
No gene flow
If at least one of these conditions is NOT met, then the population is EVOLVING
Allele frequencies: a gene with 2 alleles: p, q
p = frequency of dominant allele (A)
q = frequency of recessive allele (a)
p + q = 1
USE DECIMALS
Genotype frequencies: 3 genotypes (AA, Aa, aa)
p2 + 2pq + q2 = 1
p2 = AA (Homozygous dominant)
2pq = Aa (Heterozygous)
q2 = aa (Homozygous recessive)
If you are given the genotypes, calculate p and q by adding up the total # of A and a alleles
If you know phenotypes, then use “aa” to find q2, and then q. (p = q-1)
Use p2 + 2pq + q2 to find genotype frequencies
The key to solving H-W problems lies with the homozygous recessive individuals
Minor Causes:
Mutations: Rare, very small changes in allele frequencies
Nonrandom Mating: Affect genotypes, but not allele frequencies
Major Causes: Natural Selection, genetic drift, gene flow
Founder Effect: A few individuals isolated from larger populations
Bottleneck Effect: Sudden change in the environment drastically reduces population size
3 types of Natural Selection:
Directional Selection
Disruptive (diversifying) Selection
Stabilizing Selection
Sexual Selection: Form of natural selection - certain individuals more likely to obtain mates
Sexual Dimorphism: Difference between 2 sexes - size, color, ornamentation, behavior
Chapter 24: Speciation
Speciation: Origin of species
Microevolution: Changes within a single gene pool
Macroevolution: Evolutionary changes that cause one species to turn into 2 different species
Reproductive isolation: barriers that prevent members of 2 species from producing viable, fertile offspring
Prezygotic Barriers:
Impede Mating/Fertilization
Ex. Habitat Isolation, Temporal Isolation, Behavioral Isolation, Mechanical Isolation, Gametic Isolation
Postzygotic Barriers:
Prevent hybrid zygote from developing into viable adult
Ex. Reduced Hybrid Viability, Reduced Hybrid Fertility, Hybrid Breakdown
Allopatric Speciation: A population forms a new species while geographically isolated from its parent population
Sympatric Speciation: A subset of a population forms a new species without geographic separation
***Polyploidy means instant speciation (for plants)***
Adaptive Radiation: Many new species arise from a common ancestor
Occurs when:
A few organisms make a way to new, distant areas (allopatric speciation)
Environmental Change -> Extinctions -> New Niches for Survivors
Ex. Hawaiian Archipelago
Hybrid Zones:
Incomplete Reproductive Barriers
Possible Outcomes: reinforcement, fusion, stability
Tempo of Evolution:
Gradualism
Common Ancestor
Slow, Constant Change
Punctuated Equilibrium:
Eldridge & Gould
A long period of stasis punctuated by short bursts of significant change
Chapter 26: Cladistics/Phylogeny
Tools used to determine evolutionary relationships:
Fossils
Morphology (homologous structures)
Molecular Evidence (DNA, amino acids)(DNA does not lie)
Phylogenetic Tree: branching diagram that shows the evolutionary history of a group of organisms
***Nodes show where a common ancestor is between organisms***
Cladogram: Diagram that depicts patterns of shared characteristics among taxa
Clade: group of species that includes ancestral species + all descendants
Shared derived characteristics are used to construct cladograms
***Branch lengths can indicate time***
3 Domains: Bateria, Archea, Eukarea
***Nodes can be rotated***