Lecture 5: Sensation and Perception
Sensation: Act of using sensory systems to detect environmental stimuli
- What stimuli can our body detect, and how?
Perception: Conscious recognition and identification of a sensory stimulus
- How does brain make sense of the information our sense have detected?
Senory Receptor Cells
- Specialized cells that convert a specific form of environmental stimuli into neural impulses
o Sensory transduction: Process of converting stimuli into a neural impulse that our brain can read
Limits of the Senses
- Thresholds: Our senses require a certain amount of sensory input to detect things
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- Absolute Threshold: Smallest amount of stimulus that one can detect
o Smell: A drop of perfume diffused throughout a six-room apartment
o Taste: 5 mm of sugar in 9 litres of water
o Touch: Insect’s wing falling on your cheek from a centimeter of height
o Hearing: Tick of a watch 6 meters away in a quiet room
o Sight: Candle flame 50 km away on a clear, dark night
- Difference Threshold (just noticeable difference): Minimal difference needed to notice a difference between two stimuli
o Sensory sensitivity depends on experience, expectations, and consequences for failure
- Signal Detection Theory: Response to a signal depends on the ability to differentiate between the signal and noise, and on their response criteria
o E.g. Explores how strong a signal must be for detection
Sensory Adaptation
- Repeated stimulation of a sensory cell leads to reduced response
o Ex. Tag in your shirt bothered you in the morning, but now it is unnoticeable.
Bottom-up processing
- Perception where environmental stimuli are transformed to neural impulses that successfully move into more complex brain regions
o Ex. Light from friend’s face is converted to neural impulses that travel into the brain to visual regions
Top-down Processing
- Perception process led by cognitive processes, such as memory or expectations
o Ex. When you look at your best friend, brain regions that store information about what faces look like can help perceive and recognize visual stimulus
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The Senses
- Nervous system is composed of different sensory systems that transduce different stimuli into neural impulses
o Smell, touch, taste, sound, sight, kinesthetic (body’s movement, posture), vestibula (head’s position and movement), and thermoception (temperature detection) and nociception (pain detection)
The Chemical Senses
- Olfactory sense (smell), Gustatory sense (taste)
- Smell
o Odourants: Airborne chemicals that are detected as adours
o Olfactory receptor neurons: Cells that convert chemical signals from odourants into neural impulses
- Taste
o Papillae: Bumps on tongue that contain clumps of taste buds
o Taste buds: Clusters of sensory receptor cells that convert chemical signals from food into neural impulses
§ Includes sweet, sour, bitter, salt, and umami (taste of MSG) receptors
§ Babies like sweet more than sour, prefer odour of mother’s milk
Individual Differences in Taste and Smell
- Females are more sensitive to odours than males
- Taste is variable in individuals; some more sensitive to bitter than others
Smell and Taste Disorders
- Ageusia: Inability to taste
- Anosmia: Inability to detect odours
- Hyposmia: Reduced ability to smell
- Reflex epilepsy: Seizure occurs only after exposure to a specific odour
- Migraine headaches: Specific odours can trigger migraines
Tactile or Cutaneous Senses
- Tactile or somatosensory system: Combination of skin senses
o Pressure
o Touch
o Temperature
o Vibration
o Pain
Different Somatosensory Receptors
- Free nerve endings
o Located near surface of the skin
o Detects touch, pressure, pain, and temp
- Meissner’s corpuscles
o Located in fingertips, lips, and palms
o Transduce information about sensitive touch
- Merkel’s discs
o Located near surface of the skin
o Transduce information about light to moderate pressure against the skin
- Ruffini’s end-organs
o Deep in the skin
o Register heavy pressure and movement of joints
- Pacinian corpuscles
o Deep in skin
o Respond to vibrations and heavy pressure
Two Pathways of Pain
- Fast pathway: Sharp, localized pain is felt quicker because it travels along myelinated neurons to the brain
- Slow pathway: These inputs communicate with brain regions involved in processing emotions; pain we perceive via the slow pathway is more often burning pain than sharp pain
Development of Tactile Senses
- Developed at birth
- Ability to respond to different somatosensory stimuli is dependent on brain development and learning
Tactile Senses: Individual differences
- Women have lower threshold for detecting pain than men
- Neuroimaging studies have revealed that people’s brains react differently depending on their sensitivity to pain
Disorder of the Tactile Senses
- Chronic pain is most common abnormality
- No pain: Rare genetic condition resulting in inability to detect pain or temperature
- Phantom limb sensations: tactile hallucinations of touch, pressure, and pain in body part that no longer exists
Auditory Sense
- Sound waves: Vibrations of the air in the frequency of hearing
- Frequency: Number of cycles per second in a wave: measured in Hz
- Amplitude: Magnitude (height of a wave): measured in dB
How the Ear Hears
- Sound waves enter outer ear
- Waves hit the eardrum
- Waves pass into the middle ear
- Stapes hits the oval window, causing a wave to form in the fluid of the cochlea
- Fluid deflects the basilar membrane, bending its hair cells that transduce the sound wave into electrical activity
- As hair cells move, neural impulses are created and sent to the brain
Identifying Frequency and Pitch
- Frequency Theory: Different sound frequencies are converted into different rates of action potentials
o High-frequency sounds produce a more rapid firing
- Place Theory: Differences in sound frequency activate different regions of the basilar membrane
o Brain equates the place activity occurred on the basilar membrane with a particular frequency
Sound Adaptation
- Muscles around our ears can contract so less sound waves enter the ear
- Ears become less sensitive to continuous noise
- Our brains filter out sounds that are not important
- Cocktail party effect: Brain picks up relevant sounds even in noisy environments
Sound Localization
- General loudness: Louder sounds seem closer
- Loudness in each ear: The ear closer to the sound hears a louder noise than the other
- Timing: Sound waves will reach the ear closer to the source of the sound before they reach the ear farther away
Development of Hearing
- Babies can hear before they are born
- Ability to recognize and respond appropriately to sounds develops in first few months
- Infants prefer speech to non-speech
Hearing Loss
- Deafness: Loss of hearing
o Can be genetic, infectious, physical, exposure to toxins, etc.
- Tinnitus: Ringing in ear
o Due to abnormalities in the ear
Visual Sense
- How the eye works
o Light enters eye
o Iris adjusts pupil size to control amt of light
o Iris also changes shape of the lens to focus
o Lens focuses the light on the retina, a multi-layered sheet of nerve cells
o Photoreceptors (sensory receptor cells for vision called rods and cones) are located in retina
o Rods and cones transduce light waves into a neural impulse
The Retina
- Rods
o Detects light
o Used for periphery and night vision
o Not as acute as cones
o Many more rods than cones
- Cones
o Used for central and colour vision
o Very acute (very clear)
§ The fovea (centre of retina) contains all cones
o Less than rods
Seeing in colour
- Hue: Experience of colour based on the wavelength of light
- Saturation: Purity of color: how bright or vivid it is
- Brightness: How much light is reflected from the object
Theories for how we see colour
- Trichromatic theory: Three different snsors for colour and each type responds to a different range of wavelengths of light
- Opponent process theory: Colour pairs work to inhibit one another in the perception of colour
Visual Pathway
- Optic nerve contains the axons of 1 million ganglion cells that exist the eye via the blind spot and project to the thalamus
“What” Pathway
- Helps determine identity
- Visual agnosia: Damage to the “what” pathway; cannot recognize objects
- Prosopagnosia: A form of visual agnosia in which people cannot recognize faces
“Where” Pathway
- Locating objects in space
- Hemi-neglect: damage to the “where” pathway: people ignore one side of their visual field
o People with damage to the right side of “where” pathways neglect left side of their visual field
Gestalt (Top-Down Processing) Laws
- Proximity: Objects physically close together are grouped together
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- Similarity: Similar objects are grouped together
- Continuity: Objects that continue a pattern are grouped together
- Closure: We fill in small gaps in objects to perceive them as whole objects
- Figure ground: Tendency to perceive one aspect as the figure and other as the background
Depth Perception
- Binocular cues: Cues from both eyes
- Retinal Disparity: different images of objects are cast on the retinas of each eye
- Convergence: Tendency of eyes to move toward each other as we focus on objects up close
Monocular Cues
- Relative height: We see objects that are higher in our visual plan e as further away than those that are lower
- Texture gradient: We can see more details of textured surfaces when they are closer
- Relative Height: When two objects we know are the same size, the smaller one we recognize as further
- Linear perspective: parallel lines seem to converge in the distance
Perceptual Constancies: Shape and Size Constancy
- Perceptual constancy: Our top-down tendency to view objects as unchanging, despite shifts in the environmental stimuli we receive
- Size constancy: We perceive objects as the same size regardless of the distance from which it is viewed
- Shape constancy: we see an object as the same shape no matter from what angel it is viewed
The Visual Sense: Sight
- Development of sight
o Newborns can see, but their vision will improve significantly by two months
o At eight months, visual acuity is similar to adults
o Experience with the visual world is necessary for normal visual system development
- Visual Impairment
o Strabismus: Lack of coordinated movement of both eyes
o Amblyopia: Loss of visual abilities in a weaker eye
Other senses
- Kinesthetic – receptor cells in your muscles tell the brain when we are moving and where our body parts are in space
- Vestibular – located in the semicircular canals of our inner ears; the movement of fluid tells us if we are standing up or swaying from side to side
Lecture 7: Learning
Learning
- Lasting change caused by experience
- Associative Learning: Connections are formed between two or more stimuli; accounts for most learning
o Conditioning: The association of events in the environment
o Classical Conditioning: A form of associative learning between two previously unrelated stimuli that results in a learned response
§ Unconditioned stimulus (US): a stimulus that on its own elicits a response (food)
§ Unconditioned response: a physical response elicited by an unconditioned stimulus (does not need to be learned). i.e. salivation
§ Conditioned stimulus (CS): neutral stimulus that eventually elicits the same response as an unconditioned stimulus which it has been paired (bell)
§ Conditioned response (CR): physical response elicited by a conditioned stimulus; acquired through experience and is usually same as unconditioned response
§ Processes of Classical Conditioning
· Acquisition: Initial learning of the stimulus-response relationship
o The more pairings between CS and US, the more likely the association is learned
· Extinction: Reduction of a conditioned response after repeated presenatations of conditioned stimulus alone
· Spontaneous recovery: re-emergence of a conditioned response some time after extinction has occurred
· Stimulus generalization: What occurs when stimulis similar to the original conditioned stimulus trigger the same conditioned response (people scared of snakes are scared of all snakes, not just the first encountered one)
· Stimulus discrimination: the ability to tell the difference between one stimulus and similar stimuli
· Higher-order conditioning: What occurs when a previously conditioned stimulus functions as if it were an unconditioned stimulus for further conditioning (if pavlov started pairing music with the bell and music leads to salivating)
- Non-associative learning: Learning that does not involve forming associations between stimuli; learning occurs following repeated exposure to a single stimulus or event
o Habituation: Weaking of response to a stimulus after repeated presentation
o Dishabituation: There is a recovery of attention to a novel stimulus following habituation
o Sensitization: A strong stimulus results in an exaggerated response to the subsequent presentation of weaker stimuli
Phobias
- Persistent, irrational, or obsessive fear of a specific object or situation that may arise as a result of fear conditioning
- Systematic desensitization: process to remove phobias with gradual exposure
Operant Conditioning
- Behavior is modified depending on its consequences (Form of associative learning)
- Law of effect: Behaviours leading to rewards are more likely to occur again, while unpleasantness are less likely.
Behaviourism and Operant Conditioning
- Systematic study and manipulation of observable behaviour
- B.F. Skinner: Organisms operate on the environment
Reinforcement
- Reinforcer: Experience that produces an increase in a certain behaviour
- Positive reinforcement: Pleasant consequence following a behaviour to make it more likely to reoccur
- Negative reinforcement: Removal of an unpleasant stimulus after a response to make it more likely to reoccur
o Ex. Putting seatbelt on to remove the beeping, phobia is negatively reinforced when individual escapes anxiety by avoiding the object
- Punishment: Experience that decreases certain behaviour
- Positive Punishment: Presentating an unpleasant consequence following a behaviour
- Negative Punishment: Removal of pleasant stimulus as a consequence to a behaviour to make it less likely to reoccur
Types of reinforcers
- Primary reinforcers: A stimulus that has survival value
o Food, water, termination of pain
- Secondary reinforcers; neutral stimulus that becomes rewarding when associated with a primary reinforcer; learned
o Money, grades, praise, approval
Types of Punishers
- Primary punisher: a stimulus that is naturally aversive to an organism
o Slapping, loud sounds, extreme temperatures
- Secondary punishers: Stimulus that becomes aversive when associated with a primary
o Disapproval, criticism, bad grades
Schedules of Reinforcement
- Continuous: Reinforced every time it occurs
- Intermittent: Behaviour only followed by reinforcement some times
o Ratio schedule
§ Fixed ratio schedule: Reinforcement occurs after a fixed number of responses
§ Variable ratio schedule: Reinforcement occurs after an unpredictable, average number of responses
o Interval schedule
§ Fixed interval schedule: Reinforcement occurs every time a fixed time has elapsed
§ Variable interval schedule: Reinforcement occurs after varying lengths of time
Operant Conditioning and New Behaviours
- Shaping: Introducing new behaviour by reinforcing behaviour close to the desired behaviour
o Ex. Teaching a dog to roll over
- Behavior modification: systematic approach to change behaviour using principles of operant conditioning
Learned Helplessness
- Situations in which repeated exposure to inescapable punishment eventually produces a failure to make escape attempts
Observational Learning
- Observational learning: Occurs without overt training in response to watching models
- Vicarious learning: Occurs when an individual observes the consequences to another’s actions then chooses to duplicate the behaviour or refrain from doing so
- Mirror neurons: Neurons fired when an animal or human performs an action or when they see another perform the same action
Learning and Cognition
- Implicit learning: Refers to acquisition of information without awareness
- Spatial navigation learning: Involves formal associations among stimuli relevant to navigating in space
- Latent learning: a form fo learning that is not expressed until there is a reward or incentive
- Insight learning: a sudden realization of a solution to a problem or leap in understanding new concepts
Factors that facilitate learning
- Timing: Multiple exposures separated by time facilitate learning facts
- Context: Studying in several different locations increase slikelihood to form strong memories about the information
- Awareness and attention: Can enhance learning
Stroop Effect
- Delay in reaction time between neutral and contradicting stimuli