Classifying Organisms
Taxonomy
- Taxonomy: science of describing, naming, and classifying living and extinct organisms and viruses
- Systematics: study of biological diversity and the evolutionary relationships among organisms, both extinct and modern
- Systematists use fossil, molecular, and genetic data to infer evolutionary relationships
- Hierarchical system involving successive levels
- Taxon: each group at any level
- Domain: the highest level
- All of life belongs to one of 3 domains: bacteria, archaea, and eukarya
- Phylogenetics: reconstructing the evolutionary relationships among organisms
Cladistic Approach
- Cladistic approach: compares homologous traits, also called characters, which may exist in two or more character states
- Shared primitive character or symplesio-morphy: shared by two or more different taxa and inherited from ancestors older than their last common ancestor
- Shared derived character or synapo-morpy: shared by two or more species or taxa and has originated in their most recent common ancestor
- Basis of the cladistic approach is to analyze many shared derived characters to deduce the pathway that gave rise to those species
Constructing a Cladogram
- Choose species
- Choose characters
- Determine polarity of character states
- Primitive or derived?
- Analyze cladogram
- All species (or higher taxa) are placed on tips in the phylogenetic tree, not at branch points
- Each cladogram branch point should have a list of one or more shared derived characters that are common to all species above the branch point unless the character is later modified
- All shared derived characters appear together only once in a cladogram unless they arose independently during evolution more than once
Principle of Parsimony
- Preferred hypothesis is the one that is the simplest for all the characters and their states
- Challenge in a cladistic approach is to determine the correct polarity of events
- It may not always be obvious which traits are primitive (came earlier) and which are derived (came later in evolution)
- Fossils may be analyzed to help resolve