Week 1 - Biodiversity and History of Life

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Importance of Organismal Diversity

  • Organismal diversity is integral for ecosystem resilience, productivity, and long-term stability. Its loss can lead to fragile ecosystems that are more prone to collapse.

  • Supports ecosystem services critical for human survival such as:

    • Pollination: Essential for the reproduction of many plants/environments, which in turn supports food systems.

    • Nutrient Cycling: Critical for maintaining soil health and fertility, allowing for sustainable agriculture and natural ecosystems.

    • Climate Regulation: Diverse ecosystems help stabilize climate by absorbing carbon dioxide and regulating temperature.

  • Provides vast resources for human use including food, medicine derived from various species, and raw materials (fibers, timber).

Key Biodiversity Studies (Tilman et al.)

  • Location: Cedar Creek Ecosystem Science Reserve, Michigan.

  • Research Focus: Investigated species richness in various experimental grassland plots.

  • Key Findings:

    • Increased biodiversity correlated with higher plant biomass production, leading to more robust ecosystems.

    • Enhanced CO2 absorption efficiency observed with greater biodiversity, emphasizing the ecological value of plant variety.

  • Conclusion: Biodiversity has a direct positive impact on ecosystem productivity and stability, rendering diverse ecosystems more capable of withstanding disturbances.

Impact of Grass Species on Productivity

  • Increased diversity in grass species leads to greater primary productivity and biomass accumulation.

  • This diversity also enhances resilience against drought conditions, showcasing the benefits of polyculture systems in agriculture.

Ecosystem Services and Human Benefit

  • Studies reveal that higher biodiversity levels correlate with improved ecosystem services, directly benefitting humanity (e.g., ranging from clean air and water to pollination of crops).

Species Count

  • Approximately 1.5 - 1.75 million species have been described scientifically, but many remain unnamed or misunderstood.

  • Well-known groups: Include butterflies, flowering plants (angiosperms), and vertebrates like mammals, birds, reptiles.

  • Poorly understood groups: Encompass microbes such as bacteria, nematodes, fungi, and mites.

  • Estimates: Suggest around 8.7 million species exist on Earth, with a significant number classified as insects and plants.

Pyramid of Diversity

  • Most multicellular organisms are represented by insects, illustrating their vast diversity.

  • Significant extant species include:

    • Insects: Estimated at 8,750,000 species.

    • Plants: Approximately 320,000 known plant species.

Interpreting Data: X-Y Plots

  • Skills Check:

    • Explanatory Variable: lizard size (cm)

    • Response Variable: lizard running speed (m/s)

  • Students will learn to plot data trends from experiments involving various lizard populations to analyze relationships.

Geological Events and History of Life

Major Geological Events
  • Earth's Formation: Approximately 4.6 billion years ago (bya), initiated conditions for life.

  • Cooling Phase: Around 4.0 billion years ago saw the formation of liquid water essential for life.

  • Oxygenation Event: Major oxygenation events around 2.5 billion years ago drastically altered Earth's atmosphere and led to aerobic life forms.

  • Early Earth Conditions: Characterized by extremely high temperatures and significant volcanic activity, ultimately leading to oceans conducive for life's emergence.

Hypotheses for the Origins of Life
  • Miller-Urey Experiment:

    • Demonstrated that organic molecules (amino acids) could be synthesized under simulated early Earth conditions, supporting biogenesis theories.

  • Hydrothermal Vent Hypothesis: Suggests that life may have originated in nutrient-rich, energy-exploiting environments deep under the sea, around hydrothermal vents.

Major Events in Life's History

  • First unicellular life: Emerged approximately 3.8 billion years ago, setting the foundation for all life forms.

  • First multicellular life: Occurred around 1.2 billion years ago, indicating increased complexity in biological forms.

  • Colonization of land: Began approximately 500 million years ago, a critical step for terrestrial ecosystems.

  • Dinosaurs: Dominated the planet from 230 million to 65 million years ago, shaping ecosystems before their extinction.

  • First humans: Emerged approximately 2 million years ago, showcasing rapid development and adaptability of species in the evolution continuum.

Fossil Record and Fossilization

Fossilization Process
  • Death of Organism: Organisms with hard parts (bones, shells) have higher chances of fossilization.

  • Decay: Soft tissues decompose rapidly, favoring preservation of hard structures.

  • Burial: Rapid sedimentation enhances preservation rates.

  • Mineralization: Minerals infiltrate and replace organic material over time.

  • Compaction and Cementation: Layers harden and become rock formations through geological processes.

  • Uplift: Geological activity exposes fossils at the Earth's surface over millions of years.

Types of Fossils
  • Trace Fossils: Provide evidence of behaviors like tracks, burrows, and coprolites (fossilized feces), offering insights into past life.

Dating Fossils
  • Relative vs. Numerical Dating: Different techniques used to determine age and sequence of fossil deposition.

    • Relative Dating: Establishes a timeline of events or relative ages without specific ages.

    • Numerical Dating: Assigns numerical ages to fossils/rocks through various methods.

    • Key Dating Methods:

      • Uranium-Lead Dating: Applicable for dating rocks from 10 million years to 4.6 billion years.

      • Potassium-Argon Dating: Useful for dating specimens from 100,000 years to 4.6 billion years.

      • Carbon-14 Dating: Effective for more recent fossils (up to 100,000 years) with a half-life of 5,730 years.

Limits of Fossilization
  • Factors Influencing Fossilization:

    • Organisms with hard body parts have higher fossilization success rates.

    • Rapid entombment in sediment enhances preservation likelihood.

    • Environmental conditions such as aquatic versus arid habitats significantly impact decay processes.

Eons and Eras in Geological History

Overview
  • Precambrian: Characterized predominantly by single-celled life forms that paved the way for complex organisms.

  • Phanerozoic: An era marked by an explosion of diverse multicellular life forms, represented by animal and plant phyla.

Proterozoic Era Highlights
  • Cambrian Period:

    • Known for major diversification of animal life leading to complex ecosystems.

    • Key adaptations such as complex body plans, hard shells for protection, and efficient locomotion evolved during this period.

Mesozoic Era Highlights
  • Dominant Species: Often referred to as the Age of Reptiles, where dinosaurs thrived alongside the first birds.

  • Ended with a mass extinction event 66 million years ago that brought significant changes to the Earth's biodiversity.

Cenozoic Era Overview
  • Dubbed the "Age of Mammals," this era saw the diversification of mammals into various niches alongside flourishing bird and flowering plant populations.

  • Human evolution occurred during this era, showcasing a remarkable biological adaptation and environmental influence on species development.

Evolution by Changes in Regulation

  • Tetrapod Limb Evolution: Regulatory changes in Hox genes were crucial for the development of limbs from ancestral fin-like structures, illustrating adaptation over time in response to environmental challenges.

Visual Summary of Deep Time and Life on Earth

  • This section will examine Earth's age, environmental conditions, and the life forms evolving through time, demonstrating the intricate processes that have led to current biodiversity.

  • Key Factors:

    • Climate changes, influenced by atmospheric variations (e.g., CO2 and O2 levels), played significant roles in life evolution.

    • Fossils serve as vital records providing insights into organic evolution and the diversity of life past and present.

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