Parasitism and Mutualism
Parasitism and Mutualism Notes
Symbiosis
- Definition: Symbiosis refers to the intimate and enduring relationship between two or more organisms of different species.
- Implications: The relationship determines the fate of individuals of one species based on their interaction with another species.
- Types of Outcomes:
- Mutualism (++): Both species benefit from the relationship.
- Parasitism (+-): One species benefits at the expense of the other.
- Commensalism (+0): One species benefits while the other is unaffected.
- Amensalism (-0): One species is harmed while the other is unaffected.
Parasitoids vs. Parasites
- Parasites: Typically do not kill their hosts; they draw resources from them.
- Parasitoids: Lay eggs in a host; larvae consume the host, eventually leading to the host's death.
Characteristics of Parasites
Microparasites
- Traits: Small size, short generation time, rapid multiplication within the host.
- Associated with: Disease.
- Infection Duration: Short in relation to host lifespan, typically transmitted directly from host to host or via a carrier (e.g., viruses, bacteria).
Macroparasites
- Traits: Larger size, longer generation time, usually do not complete life cycle in a single host.
- Transmission: Can occur directly between hosts or via carriers/intermediate hosts (e.g., flatworms, roundworms, lice).
Parasitic Plants
- Over 4000 species of parasitic plants exist.
- Haustorium: A modified root that penetrates host tissues to connect with vascular tissue.
- Types:
- Hemiparasites: Photosynthetic, drawing water and nutrients from the host's xylem.
- Holoparasites: Non-photosynthetic, obtaining all resources from the host's phloem and xylem.
Host Habitats and Parasite Life Cycles
- Ectoparasites: Live on the surface of hosts (e.g., skin, feathers).
- Endoparasites: Live within hosts, potentially affecting blood and organ systems.
- Life cycles can involve multiple hosts and stages; definitive hosts are where parasites achieve sexual maturity, and intermediate hosts support lifecycle stages.
Transmission of Parasites
- Direct Transmission: Occurs between hosts without intermediate organisms (e.g., through air or water).
- Indirect Transmission: Involves an intermediate vector that transmits the parasite from one host to another.
Host Responses to Parasitism
- Behavioral Defenses: Hosts can groom to remove ectoparasites, relocate to areas with fewer parasites (e.g., deer using dense, shaded spots).
- Immune Defenses:
- Inflammatory Response: Triggers release of histamines, increased blood flow, and white blood cell activity.
- Cysts: Hard encapsulated structures around parasites to isolate them.
Impact on Host Survival/Reproduction
- Parasitism may lead to decreased fitness (e.g., smaller egg sizes in infected females).
- Altered behaviors in hosts can increase vulnerability to predators (e.g., rabbits with tularemia, killifish with trematodes).
Emerging Diseases and Human Health
- Emerging Infectious Diseases: Diseases that have recently increased in incidence or range.
- Lyme Disease Example: Transmission through ticks that obtain bacteria from infected animals, affected by human-induced environmental changes.
Mutualism
Evolution from Parasitism to Mutualism
- In cases where parasites are less harmful and actually beneficial, relationships can shift from parasitism to mutualism (e.g., tapeworms aiding in host growth).
- Type of Mutualists:
- Obligate: Cannot survive without the interaction.
- Facultative: Can survive without the interaction.
Specific Mutualistic Interactions
- Coral-Algae Symbiosis: Corals obtain most of their energy from symbiotic algae (zooxanthellae), while algae receive shelter.
- Plant-Bacteria Relationships: Legumes associate with nitrogen-fixing bacteria (Rhizobium) to receive nitrogen, while bacteria gain carbon from plants.
- Mycorrhizal Fungi: Assist plants in nutrient uptake while receiving carbon, with endomycorrhizae and ectomycorrhizae being types of interactions.
Defense and Reproduction Mutualisms
- Defense: Examples where one species helps defend another, like alkaloids from fungi in grasses that deter grazers.
- Reproduction: Pollinators like insects and birds that not only gain food but also help fertilize plants by transferring pollen; might incur energy costs to the plant.
Seed Dispersal Mutualisms
- Ants and Myrmecochore Plants: Seeds that ants collect for their elaiosome (food body) to consume, helping disperse the seeds.
- Fruit-Eating Animals: Consume fruit while ensuring seeds are not eaten, promoting germination away from the parent plant.