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Evolution on populations
Biogeography - the study of the geographic distribution of species, provides support for evolution
Microevolution - the change in allele frequencies in a population over generations, this is evolution at its smallest scale
Gene pool - all copies of every allele at every locus with all members of a population
Mutations
Most mutations are neutral or have negative effects, occasionally the consequences are beneficial
Point mutation is a change in a single base pair in DNA
Chromosomal level mutations are a change in the number or composition of chromosomes
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Evolutions of populations
Natural selection
Variation exists ang individual organisms that make up a population
Some of the trait differences are heritable
Survival and reproductive success are highly variable
Subset if individuals that survive best and produce the most offspring is not a random sample of the population
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Microevolutionary forces
Natural selection 4 postulates
Variation exists and individual organisms that make up a population
Some of the trait differences are heritable
Survival and reproductive success are highly variable
Subset if individuals that survive best and produce the most offspring is not a random sample of the population
Genetic drift
The smaller the sample the greater the chance of random deviation from a predicted result
A process in which chance events cause allele frequencies to fluctuate unpredictably from one generation to the next
Genetic drift tends to reduce genetics variation through the random loss of alleles
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Genetic drift - chance events cause unpredictable fluctuations in allele frequencies from one generation to the next
Founder effect
Occurs when a few individuals become isolated from a larger population
Allele frequencies in the smaller founder population are different from those in the parent population
ex) genetic drift could occur if a few individuals are indiscriminately blown to a new island by a storm
New population established by small group
Bottleneck effect
Occurs when there is a drastic reduction in population size due to a sudden change in the environment
The resulting gene pool may no longer be reflective of the original populations gene pool
If the population is small it may be further affected by genetic drift
Loss of significant proportion of original population
Gene flow
Consists of movement of alleles among populations
Allele can be transferred through the movement of fertile individuals or gametes ex) pollen
Tends to make populations more similar to each other over time
Speciation - forms a conceptual bridge between the processes of microevolution and macroevolution
Microevolution - consists of hinges in allele frequency in a population over time
Macroevolution - refers to broad patterns of evolutionary change above the species level
Speciation is a splitting event that results from a two step process
Genetic isolation, barrier to gene flow isolated populations
Genetic divergence, mutation, selection and genetic drift in isolated population cause divergence
Species concepts
Species - evolutionary independent population or group of populations
The biological species concept
Reproductive isolation, results in lack of gene flow between populations
Members of populations do not interbreed to fail to produce viable, fertile offspring after mating
The morphological species concept
Individual lineages differ in size shape or other morphological features
Distinguishing features most likely arise if populations are independent and isolated from gene flow
Usbale for fossils and living organisms
Can be subjective
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Monophyletic group - consists of an ancestral population plus all of its descendants also called clade or lineage
Identifies by synapomorphies
Synapomorphies - homologous traits found in a common ancestor and all the descendants but missing from more distantly related groups ex) white fur distinguishes polar bears in a monophyletic group of bears
Prezygotic barriers - block fertilization from occurring by
impending different spices from attempting to mate
preventing the successful completion of mating
hindering fertilization if mating is complete
Habitat isolation
2 species occupy different habits within the same areas may encounter each other rarely if at
Temporal isolation
Spices that breed at different times of the day, different season, or different years cannot mix their gametes
behavioral isolation
Courtship rituals and other behaviors unique to species are effective barriers to mating
mechanical isolation
Mating is attempted but morphological difference prevent its successful completion
gametic isolation
sperm of one species may not be able to fertilize eggs of another species
Post zygotic barriers - prevent hybrid zygotes from developing into viable fertile adults through
reduced hybrid viability
Genes if different parent species may interact in way that impair the hybrids development or survival in its habitat
Reduced hybrid fertility
Meiosis may fail to produce normal gametes resulting in sterility if the parent species have chromosomes of different number or structure
hybrid breakdown
First generation hybrids are viable and fertile but offspring in the next generation are feeble or sterile
mating and fertilization occur and a zygote is formed
Allopatric vs sympatric speciation
Allopatric - speciation that begins with geographic isolation. Population live in different areas are in allopatry, they live in a different homeland
Dispersal - movement of individuals form one location to another
Vicariance - physical splitting of a habitat a physical barrier splits population into subgroups that are physically isolated (river, canyon, ocean, pangea)
Geographic isolation, genetic isolation, genetic divergence
Sympatric - populations that live in the same geographic area, speciation that occurs among populations within the same geographical area
External - disruptive selection based on different ecological niches or mate preferences
Extrinsic
Intrinsic - one type of mutation polyploidy, condition of possessing more than two complete sets of chromosomes caused by massive error in meiosis or mitosis
No geographic isolation, genetic isolation, genetic divergence
Polyploidy - possessing 2 or more complete sets of chromosomes
Most plant species are the result of polyploid speciation
Plants are much more robust to abnormal chromosome numbers, as long as they can do what they need to survive they will
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Phylogeny
Speciation - the splitting of a species into 2 or more independent species
Tip - taxa are located on the tip because none of the taxa are presumed to be ancestors on others, living in present time
Branch - a line representing an evolutionary lineage, each position represents an ancestor
(node) branch point - common ancestor of dierging evolutionary lineages, most common recent ancestor, represents speciation events
Rooted tree - common ancestor of all taxa on the phylogenetic tree
Sister taxa - closely related taxa that share a recent common ancestor
Shared ancestral character- a character that originated in an ancestor of the taxon
Shared Derived character - an evolutionary novelty unique to a particular clade
Synapomorphis - found in 2 or more taxa present in their most recent common ancestor and missing in more distant ancestor
Clade (monophyletic groups) - a group that includes the common ancestor and all of the descendents, Clades are nested, like russian nesting doll there are clades within clades
Paraphyletic group - consists of an ancestral species and some but not all of the descendant, the common ancestor to all members of the group is part of the group
Polyphyletic group - group includes distantly related species but not their most recent common ancestor
Monophyletic group - a group that consists of an ancestor and all of its descendants, representing a complete branch on the tree of life, which is useful for understanding evolutionary relationships.