Litter Quality: High nitrogen, low lignin = faster.
Ecological Succession
Definition \texttt{🌿}
The natural, directional change in a community's structure and composition over time.
Occurs as ecosystems recover from disturbances or colonize new environments.
Changes During Succession \texttt{🌎}
Species Composition
Early Succession: Dominated by pioneer species (fast-growing, small, hardy organisms like grasses, mosses).
Later Stages: Slower-growing, competitive species take over (e.g., shrubs and trees).
Late Succession/Climax: Stable community with high species diversity and complex interactions.
Climax community: species replace themselves - Described by frederic clements
Biodiversity
Increases in species richness and evenness as new niches are created.
Eventually levels off or may decline slightly in mature systems.
Trends: Organic matter, moisture, and Nitrogen increase; Soil density, pH, and Phosphorous are stable then drop off
Trophic Structure
Simple food chains in early stages \longrightarrow complex food webs later.
Higher trophic levels move as habitat matures.
Biomass
Increases: more plant growth = more organic material stored in system
Soil Development
Early stages: little or no soil
Later stages: more organic material, nutrients, and structure in soil
Nutrient cycling
Becomes more efficient
Decomposition and plant uptake become faster and more balanced
Productivity
Gross and net primary production rise during early stages
May stabilize or slightly decrease in very late stages as respiration increases
Ways to Study Succession \texttt{🔎}
Chronosequence: Ordered sequences of sites with different beginning times (ex: Hawaiian Islands).
Old Field Succession:
0-15 years: non-woody plants (annual and herbaceous).
15-30 years: first woody plants (shrubs).
30-60 years: soft wood (softwood trees like tulip).
More than 60 years: hard wood (oaks).
Mechanisms of Succession \texttt{🦾}
Facilitation model: Early species improve conditions for later species.
Tolerance model: Early species are replaced by species more able to survive in new conditions (e.g., a dense, early successional tree may be replaced by a more lose/open tree).
Inhibition model: Early species prevent others from colonizing (ex: Moss dominates the intertidal zone after a disturbance)
Primary vs. Secondary Succession
Primary Succession
Definition: Succession that begins in an area with no soil or previous biological community.
Starts from: Bare rock, lava flows, glacial retreat, dunes.
Speed: Much faster than primary succession (decades to centuries) \texttt{⏱️}
Chronosequences
Definition
A chronosequence is a set of different sites that are similar in all aspects except age.
Each site represents a different stage of succession, allowing scientists to study how ecosystems change over time without waiting for decades or centuries.
Example \texttt{🔬}
Imagine four abandoned farm fields: One left alone for 1 year, one for 10 years, one for 50 years, and one for 100 years.
By comparing plant types, soil nutrients, and biomass at each site, scientists can reconstruct how succession progresses over a century.
Benefits \texttt{✅}
Saves time.
Shows long-term trends.
Reveals stages of succession.
Used in many ecosystems
Geographic Ecology
Definition
Geographic patterns of plant and animal life.
Typically LARGE scale
Biogeography: The study of regional patterns of distribution of organisms
Global Biodiversity Patterns
Diversity is lowest at the poles.
Diversity is higher at lower latitudes (tropics).
Tropics are between 23.5°N and 23.5°S.
Endemic: In one habitat and nowhere else.
Biodiversity Hotspots: High diversity.
Latitudinal Variation in Species Richness
Exceptions
Specific wasp/Insect parasitoids: Bell curve, Peak at 33°N.
Penguins: Peak is towards the south pole.
Hypotheses
Productivity: Higher primary production near the equator supports for individuals and thus more species
Environmental heterogeneity, wide variety of soil types: Diversity of habitats, More soil types, Microclimates, Provides more niches.
Niche breadth, interspecific interactions: Narrower niches and more intense interactions.
Historic and geographic factors, e.g.: time since perturbation: Less affected by disturbances.
Evolutionary rate or effective evolutionary time: Faster evolution and more time without disruption \longrightarrow speciation.
More Land \texttt{⭐}
Stable Temperature
Island Biogeography
Island Definition \texttt{🏖}
Any isolated habitat (sky islands: top of mountains or lakes: locked inside).
Have a subset of mainland species.
Equilibrium Model of Island Biogeography
Species richness is driven by:
Immigration
Extinction
Immigration Rates
Highest on a new island with no organisms (\because anything arriving is an immigrant).
As species begin to accumulate, rate of immigration declines (\because fewer arrivals would be new species).
Extinction Rates
More species = larger pool of potential extinctions.
More species = each population size is lower.
More species = more competition.
Prediction
Higher species richness on large island, near the mainland.
Landscape Ecology
Definition
Distribution and arrangement of ecosystems on a large scale.
"Studying and influencing the relationship between spatial pattern and ecological process across hierarchical levels of biological organization and different in space and time."
Mostly just about landscape structure.
Useful in conservation, land-use planning, and understanding ecological resilience.
Landscape Structure
Created by:
Geological processes
Climate
Fire
Organisms (earthworms create mounds with poop and live on the mounds)
Description of landscape structure including elements such as patches \texttt{⭐}
Patches: Discrete area that is relatively uniform compared to their surroundings
Can vary in size, shape, type
Matrix: Background or dominant land cover in a landscape that surrounds patches
Has major influence on movement and ecological flow
Corridor: Linear features that connect patches and help organisms move Describe
What differs between patches?
Biodiversity, nutrients, species richness/evenness
Methods to Capture Landscape-Scale Information
Remote Sensing
Using satellite or aircraft to gather data about the Earth’s surface.
Captures land cover types, vegetation health. Etc.
Strengths: large-scale coverage, repeated data collection
LiDAR
Sensor emits rapid laser pulses toward the ground
These pulses bounce back when they hit the surface
The time it takes for the light to return is used to calculate distances and create 3D
Satellite
Use reflective data to calculate temperature
NDVI: Use near infrared and red light to calculate vegetation
LAI: amount of leaf area per unit ground area
Ground Truthing: Verifying remote sensing data by collecting observations and measurements
GIS: store, analyze, and visualize spatial data
Puts all the layers together
Grain size (resolution) is very important
Smallest unit of measurement in spatial data
Need to choose the right resolution
Quantify patch shape based on edge-area (P/A) ratio
Perimeter Area
Edge (perimeter) to area ratio: P/A
Patch size index
Closer to 1 is circular
Edge Effect
Same area, but patch effect changes the habitat space
70% of the worlds forest are within 1 km of an edge
Land use / Land cover change
Invasive species
Human population growth
Birth rates are decreasing right now
Corridors
Connectivity
The degree to which one patch is available to individuals in another patchHow easily can species move between patches
To help this, we create corridors
How very large edge effect but
Allow for improved dispersal between patches
Tested with butter fly Both species did better in the connected patches
Conservation Biology
Definition
Study of how to conserve (protect, maintain, & restore) biodiversity.
May include aspects of population genetics, populations, community, ecosystem, geographic, and landscape ecology
Species Employed in Conservation
Indicator Species
Restricted niches
May be first lost OR only show up when there is pollution
May flies🦗; narrow niches
Umbrella Species
Require large area
If protected, also protects other species
North Atlantic Right Whale
Global Climate Change
Evidence and Causes
“Warming of the Climate System is unequivocal.” \texttt{⭐}
“It is extremely likely that human influence has been the dominant causes of the observed warming since the mid-20th century.”
Greenhouse effect:
29% reflected
23% absorbed in atmosphere
48% absorbed at the surface
Consequences
Think Pair Share
Glacier melts = sea levels rise
Extreme temperatures
Unusual seasonality
Worse hurricanes
Arrangement of patches is important
Mandelbrot (1967)
How long is the coast of Britain?
He is known for fractal geometry
Ruler Length
Climate Change Effects
Warmer on average
Extreme weather events
Droughts, hot/cold spells, heavy rain
Melting glaciers
Sea level rises
From polar ice caps
Water expanding as it gets warmer
Ocean acidification
Some CO_2 dissolved in the ocean
Challenging for agriculture \texttt{⭐}
Disease outbreaks
Temperature Changes
4-7° C change over 5000 years; Current Rate: 0.7° C last 100 years
Respiration, melting, etc. will increase rate \texttt{⭐}
Minimum temperatures will rise \texttt{☀️}
Extreme heat \texttt{🔥}
Increase in heavy rain events \texttt{🌧}
Droughts \texttt{🌵}
Species must adapt, disperse, or go extinct \texttt{⭐}
Land Use/Land Cover Change
> ½ of wetlands in the contiguous US have been drained and converted
Some ecosystem <5% of original extent remaining
Loss of tropical rainforests
Ecosystem services
Purification of air/water
Detoxification and decomposition of wastes
Cycling nutrients
Moderation of weather extremes
Ecosystem effects
Habitat loss Forest, wetlands, grasslands are cleared Reducing biodiversity
Disrupted nutrient cycle Alter nutrients like N/P
Increased invasive species disturbed/cleared land is more vulnerable to invasive species
Altered water flow Paved surfaces increase runoff and decrease groundwater recharge Affects aquatic habitats
Human Population Effects
Resources Access Improves food and housing Strain natural resources like water and soil
Health Impacts Air/water pollution from agriculture or urban expansion can lead to respiratory issues and disease