Chapter 3
Combination of all ecosystems on Earth form the Biosphere
20-km thick layer around Earth between the deepest ocean bottom and the highest mountain peak
When one organism consumes another, not all of the energy in the consumed organism is transferred to the consumer
Photosynthesis is the process of using solar energy to convert carbon dioxide and water into glucose
Glucose is a form of potential energy that can be used by a wide range of organisms
Also produces oxygen as a waste product
Solar energy + 6 H2O + 6 CO2 → C6H12O6 + 6 O2
Cellular respiration is the process by which cells unlock the energy of chemical compounds
Aerobic respiration is when cells convert glucose and oxygen into energy, CO2, and water
Anaerobic respiration is when cells convert glucose into energy in the absence of oxygen
Does not provide as much energy as aerobic respiration
Energy + 6 H2O + 6CO2 ← C6H12O6 + 6 O2
Producers both produce and consume oxygen
Generate more oxygen through photosynthesis than they consume through respiration
Overall, producers photosynthesize more than they respire
Net effect is excess of oxygen released into the air and an excess of carbon stored in the tissues of producers
Consumers are incapable of performing photosynthesis and must obtain energy by consuming other organisms
Primary Consumers
Herbivores
Include a variety of familiar plant and algae eating animals such as zebras, grasshoppers, tadpoles, and zooplankton
Secondary Consumers
Carnivores
Lions, hawks, rattlesnakes
Tertiary Consumers
Carnivores that eat secondary consumers
Bald eagles
Successive level of organisms consuming one another are knows as trophic levels
Sequence of consumption from producers through tertiary consumers is the food chain
Helps us visualize how energy and matter move between trophic levels
Food web is a complex model of how energy and matter move through trophic levels
Omnivores operate at several trophic levels
Grizzly bears who eat berries and fish
Venus Fly Trap can photosynthesize and digest insects
Each trophic level eventually produces dead individuals and waste products
Scavengers
Organisms that consume dead animals such as vultures
Detritivores
Organism such as dung beetles that specialize in breaking down dead tissues and waste products into smaller particles
Decomposers
Fungi and bacteria that complete the breakdown process by converting organic matter into small elements and molecules that can be recycled back into the system
Amount of energy available in an ecosystem determines how much life the ecosystem can support
Gross Primary Productivity
Measure of the total amount of solar energy that the producers in the system capture via photosynthesis over a given time
GPP does not subtract the energy that is lost when the the producers respire
Measure of how much photosynthesis is occurring over some amount of time
Difficult ot calculate because a plant rarely photosynthesizes and respires simultaneously
Net Primary Productivity
Energy captured by producers in an ecosystem minus the energy producers respire
Ranges from 25 to 50 percent of the GPP
Helps measure change in a system
Net primary productivity of an ecosystem establishes the rate at which biomass- the total mass of all living matter ina specific area- is produced over a given amount of time
The amount of biomass present in an ecosystem at a particular time is its standing crop
Standing crop measures the amount of energy in a system at a given time, while productivity measures the rate of energy production over a span of time
Proportion of energy that can be passed from one trophic level to another is referred to as ecological efficiency
Are fairly low: range from 5 to 20 percent
Total biomass available at a given trophic level leads to only 10 percent being converted into energy at the next higher trophic level
Trophic pyramid is representation of the distribution of biomass, numbers, of energy among trophic levels
Tend to have similar proportions across ecosystems
Most of the energy and biomass are found at the procure level and they commonly decrease as we move up the pyramid
Chapter 6
Community ecology is the study of interactions among species
Symbiotic relationships
Relationship between two species that are living in close association with each other
Include competition, predation, parasitism, and herbivory
Negative interactions
Competition
Struggle of individuals to obtain a shared limiting resource
Competitive exclusion principle
Two species competing for the same limiting resource cannot coexist
Leads to resource partitioning in which two species divide a resource based on differences in their behavior or morphology
Temporal resource partitioning
Process in which two species utilize the same resource but at different times
Spacial resource partitioning
Two species reduce competition by using different habitats
Morphological Resource partitioning
Evolution of differences in body size or shape
Predation
Interaction in which one animal typically kills and consumes the other animal
Parasitoids are animals that lay eggs inside other organisms
Defenses include behavioral, morphological, and chemical
Parasitism
Interaction in which one organism lives on or is in another organism
Single parasite rarely causes the death of its host
Pathogens are parasites that cause diseases in their hosts
Herbivory
Interaction in which an animal consumes a producer
Typically only eat a portion of a producer without killing it
Positive Interactions
Mutualism
Interaction between two species that increases the chances of survival or reproduction of both species
Under such conditions, natural selection will favor individuals that no longer engage in the mutualistic interaction
Commensalisms
Relationship between species in which one species benefits and the other species is neither harmed nor helped
Very common in nature
Keystone species is a species that is not very abundant but has a large effect on an ecological community
Beavers
Without the keystone in place, the arch would fall apart
Ecosystem engineer is a keystone species that creates or maintains habitat for other species
Beavers and alligators
Chapter 4
Terrestrial Biomes are geographic regions categorized by a particular combination of average annual temperature, annual precipitation, and distinctive plant growth forms on land
Aquatic Biomes are aquatic regions characterized by a particular combination of salinity, depth, and water flow
Tundra Biome
Cold and treeless with low growing vegetation
Growing season is usually about 4 months during the summer, when the polar region is titled toward the Sun and the days are very long
During this time, the upper layer of soil thaws, creating pools of standing water
Permafrost is an impermeable, permanently frozen layer of soil
Prevents water from draining and roots from penetrating
Permafrost and cold temperatures prevent deep rooted plants from growing
Little precipitation, but enough to support plant growth
Contains small woody shrubs, mosses, heaths, lichens
Dead plants and animals decompose very slowly
Boreal forest Biome
Forest biome made up primarily of coniferous evergreen trees that can tolerate cold winters and short growing seasons
Sometimes called Taiga
Has a very cold climate and plant growth is more constrained by temperature than by precipitation
Soil is covered in a thick layer of organic material, but poor in nutrients
Some deciduous trees such as birch, maple, and aspen can be found
Needles of coniferous trees can tolerate freezing conditions
Serve as an important source for paper,pulp, and building materials
Temperate Rainforest Biome
Coastal biome typified by moderate temperatures and high precipitation
Winters are rainy and summers are foggy
Supports growth of very large trees
Slow decomposition, but not as slow as the tundra or Boreal forest
Temperate Seasonal Forest
Biome with warm summers and cold winters with over 1 m of precipitation annually
Dominated by deciduous trees
Rapid decomposition
Have high soil fertility and support plant productivity
Said to be one of the first biomes to be converted to agriculture on a large scale
Woodland/Shrubland Biome
Biome characterized by hot, dry summers and mild, rainy winters
12 month growing season, but plant growth is constrained by high temperatures and low precipitation in the summer and by cool temperatures and high precipitation in the winter
Ideal conditions for growing grapes to make wine
Favor occurrence of wildfires
Drought resistant shrubs such as Yucca, scrub, oak, and sagebrush
Soil is low in nutrients due to leaching by the winter rains
Also called chaparral
Temperate Grassland/Cold Desert
Biome characterized by cold, harsh winters, and hot, dry summers
Plant growth is similar to Woodland
Fires are common, as the dry and frequently windy conditions fan flames ignited by lightning
Cold deserts have even sparser vegetation than shortgrass prairies
Very cold winters and do not support characteristic plan growth of hot deserts such as cacti and euphorbs
Tropical Rainforest
Warm and wet biome found between 20 degrees N and 20 degrees S of the equator, with little seasonal temperature variation and high precipitation
High productivity and rapid decomposition
High rate of decomposition causes the soils to lose their fertility quickly , so little undecomposed organic matter(humus)
Contain mot biodiversitY
Main threat is deforestation
Tropical Seasonal Forest/ Savanna
Biome marked by warm temperatures and distinct wet and dry seasons
Tropical deciduous forest
Common plants include acacia and baobab trees
Grazing and fire discourage growth of many smaller woody plants and keep the savanna landscape open
Presence of trees and warmer average annual temperature differentiate savanna from grassland
Promotes decomposition, but low precipitation constrains plants from using soil nutrients that are released
Soils are fertile
Subtropical Desert
Biome prevailing at approximately 30 degrees N and S with hot temperatures, extremely dry conditions, and sparse vegetation
Also known as hot deserts
Contains Cacti, Euphorbs, and succulent plants
When there is rainfall, growth is rapid
Aquatic Biomes are not categorized by temperature
Freshwater Biomes include streams, rivers, lakes and wetlands
Saltwater Biomes include shallow marine areas such as estuaries and coral reefs, also the ocean
Streams and Rivers
Characterized by flowing fresh water that may originate from underground springs or as runoff from rain or melting snow
Streams are typically narrow and carry relatively small amounts of water
Rivers are typically wider and carry larger amounts of water
Fast moving streams and rivers typically have stretches of turbulent water called rapids, where water and air are mixed together
The mixing allows large amounts of oxygen to enter the water and supports fish fish species
Slower moving rivers experience less mixing of air and water which favor species that can handle low-oxygen conditions
Catfish
Lakes and Ponds
Contain standing water, at least some of which is too deep to support emergent vegetation
Lakes are larger than ponds
Littoral zone is the shallow zone of soil and water in lakes and ponds where most algae and emergent plants grow
Most photosynthesis occurs in this zone
Limnetic zone is a zone of open water in lakes and ponds
Contains phytoplankton
Extends as deep as sunlight penetrates
Profundal zone is region of water where sunlight does nor each, below the limnetic zone in very deep lakes
Producers cannot survive here
Benthic zone is the muddy bottom of a lake,pond, or ocean
Lakes are classified by their level of primary productivity
Low productivity due to low amounts of nutrients such as phosphorous and nitrogen in the water are called oligotrophic
Moderate level of productivity is mesotrophic
High level of productivity is Eutrophic
Freshwater Wetlands
Aquatic biomes that are submerged or saturated by water for at least part of each year, but shallow enough to support emergent vegetation
Support species of plants that are specialized to live in submerged or saturated oils
Include swamps, marshes, and bogs
Swamps are wetlands that contain emergent tres
Marshes are wetlands that contain primarily non woody vegetation
Bogs are very acidic
Wetlands can take in large amounts of rainwater and release it slowly into the groundwater or into nearby streams
Marine biomes contain salt water and can be categorized as salt marshes, mangrove swamps, intertidal zones, coral reefs and the open ocean.
Salt Marshes
Marsh containing nonwoody emergent vegetation, found along the coast in temperate climates
Found within an estuary which is an area along the coast where the fresh water of a river mixes with salt water from the ocean
Habitat for spawning fish and shellfish
Mangrove swamps
Swamp that occurs along tropical and subtropical coasts, and contains salt-tolerant trees with roots submerged in water
Mangrove trees are salt tolerant
Intertidal Zones
Narrow band of coastline between the levels of high tide and low tide
Range from steep rocky areas to broad sloping mudflats
Coral Reefs
Most diverse marine biome on Earth, found in warm, shallow waters beyond the shoreline
Corals are tiny animals that secrete a layer of limestone to form an external skeleton
Corals live in water that is relatively poor in nutrients and food
Corals die and their skeletons accumulate to become a coral reef
Coral Bleaching is a phenomenon in which algae inside corals die, causing to turn white
Open Ocean
Deep ocean water, located away from the shoreline where sunlight can no longer reach the ocean bottom
Photic zone is the upper layer of the ocean water in the ocean that receives enough sunlight fir photosynthesis
Chemosynthesis is process used by some bacteria in the ocean to generate energy with methane and hydrogen sulfide
Biogeochemical Cycle
Movements of matter within and between ecosystems
Components that contain matter(air, water, and organisms) are referred to as pools
Processes that move matter between pools are knows as flows
The movement of water through the biosphere is known as the Hydrologic cycle
The release of water from leaves during photosynthesis is transpiration
Combined amount of evaporation and transpiration is called evapotranspiration
Helps move elements that are dissolved in the water
Harvesting trees can reduce evapotranspiration
Runoff and percolation can increase
Leads to erosion and flooding
Carbon makes up 20% of an organisms body weight68
Carbon Cycle
Movement of carbon around the biosphere
Photosynthesis, respiration, exchange, sedimentation, burial, extinction and combustion
The amount of carbon released from the ocean into the atmosphere roughly equals the amount of atmospheric CO2 that diffuses into ocean water
Portion of Carbon dioxide dissolved into the ocean combined with calcium ions to form calcium carbonate
Compound which can precipitate out of the water and form limestone and dolomite rock via sedimentation and burial
Although sedimentation is a long process, small amounts of calcium carbonate sediment formed each year have accumulated over millions of years to form the largest carbon pool in the slow part of the carbon cycle
Extraction is a relatively recent phenomenon
Extraction AND combustion alter the Carbon Cycle
Combustion, Respiration, and Decomposition all cause organic molecules to be broken down to produce CO2, water, and energy
However, respiration and decomposition are biotic processes, whereas combustion is an abiotic process
Exchange of carbon between the Earth’s surface and atmosphere is in a steady state
Excess CO2 in the atmosphere acts to increase the retention of heat energy in the biosphere
Results are global warming
Nitrogen Cycle
Macronutrients
Six key elements needed by organisms in a relatively large amount
Nitrogen, phosphorous, calcium, magnesium, and sulfur
Nitrogen is considered a limiting nutrient for producers
Nutrient required for the growth of an organism but available in a lower quantity than other nutrients
Nitrogen Cycle is the movement of nitrogen around the biosphere
Includes: Nitrogen Fixation, Nitrification, Assimilation, Mineralization, and denitrification
Nitrogen Fixation
Process that converts nitrogen gas in the atmosphere(N2) into forms of nitrogen that producers can use
Can office through biotic or abiotic processes
Biotic Fixation
Cyanobacteria can convert N2 gas into ammonia (NH3), which is rapidly converted to ammonium (NH4+), a form that producers can use
Abiotic fixation
N2 can be fixed in the atmosphere by combustion processes, which then convert N2 into Nitrate(NO3-)
Nitrate is carried to Earth’s surface via precipitation
Nitrification
Conversion of ammonium into nitrite, then into nitrate
Conducted by specialized species of bacteria
Nitrite is not used by most producers, but nitrate is
Assimilation
Process by which producers incorporate elements into their tissues
Usually when primary consumers feed on producers, producers nitrogen is assimilated into the tissue of the consumers, while rest is eliminated as waste products
Mineralization
Process by which fungal and bacterial decomposers break down the organic matter found in dead bodies and waste products and convert it into inorganic compounds
Sometimes called ammonification
Process by which fungal and bacterial decomposers break down the organic nitrogen found in dead bodes and waste products and convert it into inorganic ammonium
Denitrification
Final step which involves the conversion of nitrate in a series of steps into the gases nitrous oxide(n2o),and eventually, nitrogen has which is emitted into the atmosphere
Conducted by bacteria that live under anaerobic conditions
The Phosphorus Cycle
Movement of phosphorous in the biosphere
Primarily operates between land and water; There is no gas phase
Phosphorous rarely changes form: Typically found in the form Phosphate
Producers on land and in the water take up inorganic phosphate and assimilate the phosphorous into their tissues as organic phosphorous
Abiotic process involves movement between water and land
Phosphorous is not soluble in water, so much of it precipitates out of solution in the form of phosphate-laden sediments
Over time, geologic forces can lift these ocean layers and they become mountains
The phosphate rocks in the mountain are slowly weathered, brining the phosphorous to terrestrial habitats
Limiting nutrient in aquatic habitats
Increased phosphate in aquatic ecosystems can cause algal blooms
Water will become hypoxic: lox in oxygen
Dead zone os when oxygen concentrations become so low that it kills other aquatic animals
The Sulfur Cycle
Producers absorb sulfur through their roots in the form of sulfate ions
Volcanic eruptions are natural source of atmospheric sulfur int he form of sulfur dioxide
SO2(atmospheric sulfur) turns into sulfuric acid when it mixes with water
Carried back to earth when it rains or snow