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Chapter 5: The Perception of Color

  • Review

    • Light is both a particle(photon) and a wave

      • Reflected: energy that is redirected when it strikes a surface, usually back to its points of origin

      • Most of the light is reflected

        • typically white reflects almost all the light

    Color

    • Color DOES NOT exist in the physical world

    • Color is not a physical property but a psychophysical property

      • not being able to see color would impair survival

    • Color is how brains interpret and differentiate light information

      • color only exists in our brain

    • Most mammals have 2 cone types, humans have 3

Color Perception

“The perception of color is generated Internally”

  • Detections: The wavelength of light must be detected in the first place

    • There are 3 types of cones with different sensitivities to any given wavelength of light given

      • S-cones detect short wavelengths (“blue” range).

      • M-cones detect medium wavelengths (“green” range).

        • most sensitive to mid-wavelength

        • acquiring most information from this region of the visible spectrum

        • We can differentiate different shades of green

      • L-cones detect long wavelengths (“red” range). peak!!

      Photopic: light intensities that are bright enough to stimulate the cone receptors and enough to “saturate” the rods to their maximum response

      • Ex: sunlight and bright indoor lighting

      Scotopic: light intensities that are bright enough to stimulate the rod receptors but too dim to stimulate the cone receptors

      • Ex: Moonlight and dim indoor lighting

      • 4 types of photoreceptors give 3 color discrimination

      • rods do not contribute to color perception

        • Duplex Vision System: the idea that our visual system can operate in fundamentally different ways, depending upon the conditions in the environment

        • Scopic deals with light and core


  • Discrimination: we must be able to tell the difference between one wavelength ( or mixture of wavelengths_ and another

  • Important!! The principle of univariance is an infinite set of different wavelength and intensity combinations that can elicit exactly the same response from a single type of photoreceptor

    • Therefore, one type of photoreceptor cannot make color discriminations based on wavelength.

    • they are eliciting the same firing rate

    • receptor response is fixed whereas wavelength intensity can elicit different responses

  • Rods are sensitive to scotopic light levels

    • all rods contain the same photopigment molecules: rhodopsin

    • Therefore, all rods have the same sensitivity to different wavelengths of light

    • There is only one type of Rod

    • every rod will react in the same type of way

    • .– Consequently, rods obey the principle of univariance and cannot sense color differences.

    • There are individual differences in Cones and rod distribution which could affect periphery visions

      • lighting conditions

      • distance

      • spatial frequency

        • mid level SF is where we see best because it lines up best with our ganglion cells

      • Rate of Movement

Color Discrimination

  • Important!! Trichromacy is the theory that the color of any light is defined in our visual system by the relationships of three numbers, the outputs of 3 receptor types now known to be 3 cones.

    • Thomas Young and Hermann von Helmholtz proposed the trichromatic theory of color perception in humans aka “Young-Helmholtz Theory”

  • Trichromatic Theory is all colors can be made by matching 3 basic colors (red, green, blue)

    • we have 3 basic cone types are sensitive to corresponding wavelengths

    • degree of activation dictates color

      • ex. S-cones moat Activated, see blue

  • Metamers a different mixtures of wavelengths that look identical; generally, any pair of stimuli that are perceived as identical in spite of physical differences

  • James Maxel developed a color-matching technique used today still

  • There are 2 types of Color-Mixing

    • Additive color Mixing: a mixture of lights

    • If light A and light B are both reflected from a surface to the eye, in the perception of color, the effects of those 2 lights add together

    • Subtractive color Mixing: A mixture of pigments

      • if pigment A and B mix, some of the light shining on the surface will be subtracted by A and some by B.

        • Only the remainder contributes to the perception of color

      • subtractive color Mixing of pigments

  • Lateral Geniculate Nucleus (LGN) has cells that are maximally stimulated by spots of light.

    • visual pathways stop in LGN on the way from the retina to the visual cortex

    • LGN cells have receptive fields with center-surround organization

  • Cone-opponent cell: a neuron whose output is based on a difference between sets of cones.

    • In LGN there are cone-opponent cells with center-srround organization.

    • There are color-opponent ganglion cells that are antagonist red-green or blue-yellow centre surround


Color Appearance

  • Appearance: we want to assign perceived colors to lights and surfaces in the world and have those perceived colors be stable over time, regardless of different lighting conditions.

  • Color Space: a 3-dimensional space that describes all color

    • RGB color space: defined by the outputs of long, medium, and short wavelengths lights (ie. red, green, and blue)

    • HBS color space: defined by hue, saturation, and brightness !!Important!!

      • hue: the chromatic (color) aspect of light

      • saturation: the chromatic strength of hue

      • Brightness: the distance from black in color space.

  • Nonspectral colors: some colors that do not correspond to a single wavelength of light Ex. Purple_magneta

    • no single wavelength on the spectrum will have the hue of this mixture of long and short wavelength

  • Ewald Hering noticed that some color combinations are “legal” while others are “illegal”

  • The concept of legal and illegal colors comes from opponent process theory and the way our visual system processes color information.

  • It refers to color combinations that are either perceptually possible (legal colors) or impossible to see (illegal colors) due to the way our opponent neurons encode color information.

    • we can have bluish green (cyan, reddish-yellow(orange), or bluish red (purple)

    • but we cannot have green or bluish-yellow

  • Opponent color theory: The theory that perception of color depends on the output of three mechanisms, each of them based on an opponency between two colors: red–green, blue–yellow, and black–white.

    • Some LGN cells are excited by L-cone activation in center, inhibited by M-cone activation in their surround (and vice versa) Ex.Red versus green

    • Other cells are excited by S-cone activation in center, inhibited by(L + M)-cone activation in their surround (and vice versa).

  • Hue Cancellation experiments are used to determine the wavelength of unique hues

  • Unique hue: any of 4 colors that can be described with only a single color term: red, yellow, green, blue.

  • Afterimages: A visual image seen after a stimulus has been removed.

  • Negative afterimage: An afterimage whose polarity is the opposite of the original stimulus.

    • –Light stimuli produce dark negative afterimages.

    • Colors are complementary. Red produces green afterimages and blue produces yellow afterimages (and vice versa).

    • This is a way to see opponent colors in action.69


Steps in Color Perception

  1. Detection

    1. S-, M-, and L-cones detect light.

    2. each cone responds to a different range of wavelength light.

  2. Discrimination

    1. Cone-opponent mechanisms discriminate wavelengths.

    2. [L – M] and [M – L] compute something like red vs. green.

    3. [L + M] – S and S – [L + M] compute something like blue vs. yellow.

  3. Appearance

    1. Further transformations of the signals create final color-opponent appearance.73


Color in The Visual Cortex

  • V!, V2, and V4 all involved in color perception, but not exclusively

    • no particular place in the cortex specialized for color

    • Achromatopsia: loss of color vision from brain damage

  • Language and Color

    • General agreement on colors

    • Basic color terms: Single words that describe colors, are used with high frequency and have meanings that are agreed upon by speakers of a language.

    • Cultural relativism: in sensation and perception, the idea that basic perceptual

  • Genetic Differences in Color Perception

    • About 8% of males and 0.5% of females have some form of color vision deficiency: “color blindness.”

    • Color-anomalous: A term for what is usually called “color blindness.” Most“color-blind” individuals can still make discriminations based on wavelength. Those discriminations are just different from the norm.

    • Several types of color-anomalous people

      • Deuteranope: Due to the absence of M-cones.

      • Protanope: Due to the absence of L-cones.

      • Tritanope: Due to the absence of S-cones.

  • Cone monochromat: Has only one cone type; truly color-blind.

  • Rod monochromat: Has no cones of any type; truly color-blind and very visually impaired in bright light.

  • Anomia: Inability to name objects or colors in spite of the ability to see and recognize them. Typically due to brain damage.

    • Color anomia: Can see color but cannot name it

!!Important!!Color constancy: The tendency of a surface to appear the same color under a fairly wide range of illuminants.

Illuminant: The light that illuminates a surface.

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