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Introduction to Cellular Organization
- Multicellular organisms consist of various cell types, not just one.
- Cells organize into tissues, which are essential for function.
Types of Tissues in Animals
- Four major tissues in animals:
- Nervous Tissue: Involved in receiving and transmitting nerve impulses.
- Muscle Tissue: Responsible for movement, including smooth muscle in guts.
- Epithelial Tissue: Covers body surfaces and lines cavities.
- Connective Tissue: Supports and protects other tissues.
- Tissues consist of cells and extracellular matrix (ECM).
- Cells secrete molecules that compose ECM, enabling connections between cells.
Plant vs. Animal Cells
- Plants and animals evolved multicellular structures independently due to differing needs:
- Animals: Mobile, requiring strong and adaptable tissue.
- Plants: Stationary, requiring rigid yet flexible tissue for support.
Plant Cell Walls
- Cell walls surround, protect, and define plant cell shapes.
- Considered a form of extracellular matrix.
- Plant cells can synthesize and secrete components affecting cell wall characteristics (thick/thin).
- Cell walls are tough but can be fragile due to sensitive cell membranes.
- Lacking intermediate filaments, plant cells depend on osmotic pressure.
- Loss of water leads to plant wilting.
Composition of Cell Walls
- Primary Cell Wall: Formed during initial cell growth; can expand as cells grow.
- Secondary Cell Wall: Formed when cells stop growing; results from thickening of primary wall or adding layers.
- Different plant cells can have unique secondary wall structures.
Observation of Cell Walls
- Cross sections show primary walls (roots) versus secondary walls (stems).
- Staining methods reveal components such as pectin (primary wall) and cellulose (secondary wall).
Strength and Structure of Cell Walls
- Extracellular matrices provide tensile strength via long fibers aligning with stress lines.
- Cellulose microfibrils resist compression and tension, formed from long unbranched chains of glucose.
- Orientation of cellulose microfibrils determines the growth direction of cells.
- Cells must overcome microfibril resistance to expand in specific directions.
Growth Dynamics
- Turgor pressure facilitates cell growth, influencing direction based on microfibril orientation.
- Enzyme complexes on the plasma membrane elongate microfibrils, creating new cellulose.
- While most ECM molecules are made intracellular, cellulose synthesis is unique.
- Microtubules guide cellulose synthesizing proteins, aligning with interior cell structure; microtubules control microfibril orientation.
Summary of Cell Wall Composition
- The mechanism of cellulose production illustrates a direct link between intracellular processes and extracellular components.
- Microtubules inside the cell shape how cellulose forms in the extracellular space, directly influencing plant growth and stability.