Translation:

The Rules for the Translation of RNA into Amino Acids Are Contained in the Genetic

↳ Genetic Code → Which amino acid will be used to build a protein

↳ Codon:A sequence of three nucleotide bases that specifies one amino acid

General Map of Translation:

1. 2. The mRNA transcript is organized into codons…

Which is recognized by tRNA

↳ Has amino acid specificity,

includes an Anticodon (denoted by

the three yellow bases in the image to

the right) which is complementary to

the mRNA codon, and noncovalently

interacts with ribosomes.

↳ Each tRNA carrying an

amino acid called “charging” which

requires ATP

(denoted by pink section at the top of

tRNA)

Practice Question: The anticodon of a particular tRNA molecule is:

a. b. Complementary to the corresponding mRNA codon

The same sequence as the mRNA codon → This is incorrect because it’s not the same,

the anticodon sequence is complementary

c. The part of tRNA that binds to the specific amino acid

3. This is all occurring through interactions with

ribosomes

↳ Ribosome holds mRNA and tRNA in correct

position to assemble a polypeptide

↳ The Translation sequence occurs at 3 Sites:

E, P, and A

→ The Steps of Translation:

1. Initiation: A complex forms around the mRNA

a. The small ribosomal subunit

b. Charged (carrying an amino acid) tRNA

-

The first amino acid in a polypeptide

is always methionine (sometimes

taken out later)

c. The large ribosomal subunit

2. Elongation:During elongation, amino acids are sequentially added to the growing polypeptide chain as the ribosome moves along the mRNA strand.

a. b. c. After the initial tRNA enters the P site in

Initiation, the next codon is “open” for another

charged tRNA in the A site

The large subunit catalyzes a two-step reaction:

i. The bond between the tRNA and its amino

acid in the P site is broken

ii. A peptide bond forms between that amino

acid and the amino acid attached to the

tRNA in the A site

Therefore, the growing peptide is now in the A

Site

d. Our ribosome complex will then shift:

To recap: The next open codon is in the A Site,

The growing peptide is in the P site, the original

tRNA (without its amino acid now) is now in

the E site

e. The next tRNA will then enter our previously

open A Site

-

The growing peptide chain is

transferred to the next amino acid

-

The tRNA in the E Site (which is

uncharged) dissoassociates to become

recharged

-

The ribosome shifts again, leaving the

A Site open again

3. Termination: Occurs when the Stop Codon is in the

A Site

-

-

The Stop Codon binds a release factor which

releases the tRNA from its polypeptide chain in

the P Site

The polypeptide separates and folds into its 3D

Shape

→ What happens to proteins after the translation process?

-

They have specified functions so they have to go to the right place

↳ A Signal Sequence on the protein indicates where the protein’s “job” is

-

Depending on this sequence, translation could be moved to:

-

the RER because the proteins ultimate job is in the endomembrane system

or exported from the cell

-

OR translation could occur in the cytoplasm to allow entrance to the

nucleus

-

Post Translational Modification

↳ Phosphorylation (addition of phosphate groups), Glycosylation (addition of sugars),

Proteolysis (polypeptide cut by proteases to remove a signal sequence)