Cooperation

• Altruism: behavior that has a fitness cost to the individual exhibiting the behavior/ fitness benefit to recipient

• Seemingly paradoxical to ultimate causation

• What is kin selection?

Hamilton’s Rule

• Created a model to assess how an allele for altruistic behavior could increase in frequency

• Altruistic behavior is most likely to occur if:
  Fitness benefits for recipient are high (high B)
  The giver and taker are closely related (high r)
  The fitness costs for the