Chapter 13: Evidence of Evolution - Vocabulary Flashcards
Overview
- Chapter 13: Evidence of Evolution
- Topic focus: how we test and document evolution using multiple lines of evidence
- Learning objectives (Ch. 13, sections 13.1 - 13.6):
- Explain what different lines of evidence can tell us about evolution
- Explain how the fossil record and isotopes can shed light on the characteristics of extinct life and contribute to inference of evolution and ecology
- Explain how we use biogeography, homologies, vestigial structures and convergent evolution to document evolutionary change and possible lineage relationships
- Core idea: Species we see today are snapshots of an ongoing evolutionary process; life on Earth began long ago and has changed through time under geological influence and ecological pressures
- Key framing: the geologic timescale is used to organize Earth history around biological and geographical events
The Evolutionary Process: A Snapshot View
- Concept: Species today are not static; they represent moments in a long, branching process of evolution
- Visual takeaway: Evolutionary history is a continuum, with surviving lineages and extinct lineages contributing to present diversity
The Geologic Context and Origin of Life
- Life on Earth arose approximately 3.8 ext{ billion years ago}
- Scientists use the geologic timescale to divide Earth’s history into eons and eras, based on evidence of biological and geographical events
- Evolutionary events are studied in the context of this geologic timescale, linking biology with Earth history
Fossil Evidence and Paleontology
- Paleontology is the study of fossil remains or other clues to past life
- Fossils provide the original evidence for evolution; they reveal what ancient organisms looked like and, in some cases, how they behaved
- Fossils show both form and function (evidence of behavior through trace fossils, trackways, etc.)
- Fossils in Colorado have yielded large surface deposits (scrapes up to 2 meters in diameter) illustrating paleoenvironments
- Fossils help infer lineage relationships and ecological contexts across deep time
- Fossils form in multiple ways:
- Compression and petrification (permineralization) — often preserves hard parts in stone-like form
- Impressions and casting — external mold of an organism or internal cast filling a mold
- Intact preservation — rare, when organisms are buried rapidly in low-oxygen conditions, minimizing decay and scavenging
- Example sources and figures referenced: Section 13.2, Figure 13.4; images credit for specimens and contexts (various sources)
- Illustrated examples include: ancient remains trapped in amber, dinosaur feathers preserved, and micro-contexts like preserved trace materials
- Important rationale: The mode of preservation affects what information is recoverable (external shape, internal structures, soft tissues rarely preserved)
Fossilization in Practice: Conditions and Manifestations
- Intact preservation occurs when burial is rapid and oxygen is scarce, limiting decomposition and scavenging
- Fossilization can capture details such as skin impressions, feather imprints, and even preserved soft tissues rarely; most fossils are of hard parts
- Notable examples mentioned include: snakefly preserved in Baltic amber (~34–48 million years ago), dinosaur feathers preserved in Myanmar (~160 million years ago), and other preserved specimens
- Fossils also reveal behavioral clues through associations and arrangements that suggest social or mating behaviors (e.g., dinosaur–bird-like displays)
- Fossils in Colorado illustrate large-scale trace evidence (scrapes up to ~2 meters in diameter), indicating behavior or habitation patterns
Fossil History and Its Incompleteness
- Fossil history is incomplete for several reasons:
- Many organisms have soft bodies that fossilize poorly or not at all
- Erosion, fossil burial, and plate tectonics can destroy or relocate fossils over time
- Oklahoma marine fossils indicate historical ocean coverage in areas that are now terrestrial, illustrating shifts in environments through time
- The incomplete nature of the fossil record is a recognized limitation in reconstructing full evolutionary histories
Ideal Sequence for Fossilization (The Path to Becoming a Fossil)
- The typical sequence proposed for fossilization involves:
1) Die in a promising location
2) Avoid being consumed
3) Become buried by sediment
4) Survive through time
5) Be exposed and then found (before eroded) - Note: Most organisms never fossilize due to various ecological and taphonomic factors; fossil records reflect a biased sample of past life
Dating Fossils: Timing the History of Life
- Dating methods yield clues about when organisms lived and changed
- Relative dating:
- Dates fossils according to the rock layers in which they are found
- Assumes deeper layers are older than those above them; indirect and less precise, but valuable
- Absolute dating (radiometric dating):
- Dates the fossil using the chemistry of surrounding materials or the fossil itself
- Example method: radiometric dating by measuring the amount of ^{14}\mathrm{C} in a fossil to estimate time since death
- The combination of relative and absolute dating helps place fossils within the geologic timescale and infer rates of evolution
Isotopes and Diets: What Carbon Isotopes Tell Us
- Carbon isotopic signatures are used to infer diets of extinct species
- Isotopic analysis can reveal trophic level, feeding strategies, and ecological niches of ancient organisms
- This evidence complements morphological data to reconstruct past ecologies
Biogeography: Geography, Plate Tectonics, and Evolution
- Biogeography examines how species are distributed geographically and how geography shapes evolution
- Earth’s geography has changed drastically over the last 200 million years; plate tectonics drives continental drift and ocean opening/closing
- The theory of plate tectonics posits that forces within the planet move Earth’s land masses
- Continents continue to move today; earthquakes and volcanoes attest to ongoing plate motion
- Fossil and species distributions around 200 million years ago suggest the existence of a supercontinent named Pangaea
- Biogeography helps explain historical connections and separations of lineages across oceans and landmasses
- Wallace’s Line (a biogeographic boundary) illustrates long-term separation of fauna on either side, leading to distinct, independently evolving communities
- These geographic patterns provide evidence for historical isolation and divergence among lineages
Anatomical Evidence: Shared Body Plans and Common Descent
- Investigators examine anatomical features to infer evolutionary relationships between organisms
- Homology: features derived from a common ancestor
- Example: forelimbs of diverse vertebrates contain similar bone arrangements, suggesting a common ancestral limb structure
- Key figure: Section 13.3, Figure 13.10 illustrates anatomical homology
- Vestigial structures: reduced or unused features that are homologous to functional structures in related species
- Examples: vestigial eyes in some blind mole species; hindlimbs in some snakes; pelvises in whales
- These serve as evidence of evolutionary ancestry despite current function loss
- Analogous structures: superficially similar structures that arise independently and are not from a common ancestor
- Example: wings in birds and insects both enable flight but have different underlying structures
- Section 13.3, Figure 13.12 highlights these differences
- Convergent evolution: the process by which similar structures evolve independently in distantly related lineages due to similar selective pressures
- Developmental homology: embryonic development often reveals deeper similarities than adults
- Example: human and chimp skull development show greater similarity in fetuses than in adults
- Section 13.5, Figure 13.14 demonstrates developmental homology
- Early vertebrate embryos appear alike; adult forms diverge, making embryonic comparisons a powerful tool for inferring relationships
- Example comparisons include chick, mouse, and human embryos (illustrations referenced in figure 13.15)
Molecular Evidence: DNA, Proteins, and the Language of Life
- Molecular data provide extremely detailed insights into relatedness beyond morphology
- Sequence similarities are inherited from common ancestors; two unrelated species are unlikely to evolve exactly the same DNA and protein sequences by chance
- Section 13.6, Figure 8.4 introduces molecular homology concepts
- Changes in DNA underlie evolutionary changes; gene-level events can drive adaptation
- Example: amylase gene duplication associated with a high-starch diet shows how gene copy number changes can respond to ecological opportunities
- Example image references: chimp and human contexts for amylase gene study
- Homologous protein sequences demonstrate common descent across taxa
- Cytochrome c is a mitochondrial protein found in all eukaryotes; the degree of amino acid differences between species reflects the distance from their common ancestor
- Section 13.6, Figure 13.19 illustrates molecular homology using cytochrome c
- Molecular clocks: mutations accumulate in DNA at a relatively constant rate over time when not strongly selected for or against
- This underpins the concept that genetic differences can be translated into approximate divergence times
- Section 13.6, Figure 13.20 illustrates the molecular clock concept
Integrating Evidence: How the Pieces Fit
- Consilience of evidence across geology, paleontology, geography, anatomy, development, and molecular biology strengthens inferences about evolution and lineage relationships
- Fossil, isotopic, geographic, and molecular data collectively illuminate the history of life, the timing of major transitions, and the emergence of biodiversity
- Ethical, philosophical, and practical implications: understanding deep time informs debates about human origins, our relationship to ecosystems, and the responsibility to study and conserve life on Earth
- Section 13.1: Intro to evidence of evolution; link to geology and fossils
- Section 13.2: Fossil formation, fossilization processes, dating fossils; Figure 13.3, Figure 13.4
- Section 13.3: Biogeography and anatomical evidence; Figures 13.7, 13.8, 13.9, 13.10, 13.11, 13.12, 13.13
- Section 13.5: Developmental homology; Figure 13.14, 13.15
- Section 13.6: Molecular evidence and clocks; Figures 8.4, 13.18, 13.19, 13.20
- Notable empirical references mentioned: Lockley et al., Scientific Reports 2016 (dinosaurs dancing to woo mates); Greg Dale/Getty Images (fossil depiction); Terry Moore/Stocktrek Images/Getty Images (dating fossils); various image credits for fossils and embryos
Quick Recap: Core Concepts to Remember
- The fossil record provides direct though incomplete evidence of past life and evolutionary transitions
- Dating fossils uses relative layering information and absolute radiometric methods (e.g., dating via ^{14}\mathrm{C})
- Isotopes, especially carbon isotopes, reveal past diets and ecological interactions
- Biogeography links distributions to historical geologic events (plate tectonics, Pangaea, Wallace’s Line)
- Homology (shared ancestry) vs. analogy (convergence) vs. vestigial structures clarifies relationships and evolutionary paths
- Embryology often reveals deeper homologies that are not obvious in adults
- Molecular data (DNA/proteins) provide high-resolution evidence for relatedness and timing via molecular clocks
- The integration of multiple lines of evidence yields robust inferences about how evolution operates and how lineages are related