Chapter 22 -Lecture
Chapter 22 Geographic Ecology
22.1 Species Richness and Geographic Patterns
Species Area Relationship: Species richness increases with area and decreases with isolation.
Dynamic Immigration-Extinction Balance: Species richness can be modeled as a balance between immigration and extinction.
Latitudinal Patterns: Species richness generally increases from middle and high latitudes toward the equator.
Influence of Historical Processes: Long-term historical and regional influences affect species richness and biodiversity.
Introduction to Geographic Ecology
Definition: Geographic ecology seeks to map the patterns of plant and animal life.
Focus Areas: Primarily concentrates on island biogeography, latitudinal patterns in species richness, and the influence of historical processes on biodiversity.
22.1 Area and Isolation Effects on Species Richness
Early Contributions: Arrhenius quantitated the relationship between area and species number; a foundation for geographic ecology.
Island Area and Species Richness
Preston's Findings: Noted that the fewest bird species exist on the smallest islands, while the largest islands host the most species.
Nilsson et al.: Identified island area as a key predictor of species richness among various taxa in Sweden.
Habitat Fragmentation
Montane Islands: Following warming at the end of the Pleistocene, forests and habitats became fragmented into isolated mountain peaks, impacting both plant and animal distribution.
Lake Islands: Lakes function as islands with species diversity correlating with lake area studied by Tonn and Magnuson, as well as global studies by Barbour and Brown.
Marine Islands
MacArthur and Wilson's Findings: Isolation leads to reduced bird diversity on Pacific Islands, with observed increases in diversity with larger island sizes.
22.2 Equilibrium Model of Island Biogeography
Concept Overview: Introduced by MacArthur and Wilson; species diversity reflects a balance between immigration and extinction rates.
Immigration Rates: Highest on new islands; declines as species accumulate.
Extinction Rates: Increase with more species as competition intensifies and species populations diminish.
Prediction of Species Number: Point of intersection of immigration and extinction rates predicts the expected number of species on an island, influenced by both island size and distance from source populations.
Species Turnover
Dynamic Composition: Species composition changes over time, known as species turnover; Diamond's studies on California Channel Islands illustrated this.
Experimental Studies
Simberloff and Wilson: Investigated recolonization patterns on defaunated islands, affirming the dynamic nature of species composition.
22.3 Latitudinal Gradients in Species Richness
General Trend: Species richness increases as one moves from higher latitudes to the equator.
Hypotheses: Six categories of mechanisms explaining this pattern:
Time Since Perturbation: Tropics have older ecosystems with less disturbance, allowing for higher speciation rates.
Productivity: High energy in the tropics supports larger populations and reduces extinction risks.
Environmental Heterogeneity: Increased habitat complexity in the tropics supports diverse species.
Favorable Climatic Conditions: Less temperature variation and physically extreme environments in the tropics support greater biodiversity.
Niche Breadths and Interactions: Tropical species may be more limited by biological factors than physical.
Speciation and Extinction Rates: The tropics experience higher rates of speciation and lower extinction rates compared to higher latitudes.
Area's Role in Species Richness
Regional Area Influence: Larger land areas in the tropics allow for more habitat diversity, which supports greater species richness.
22.4 Historical and Regional Influences on Species Richness
Exceptional Patterns: Variance in species richness that cannot simply be explained by area, illustrated by the Cape region of South Africa having more species than larger areas like California.
Historic Factors: Influences such as continental drift and climate changes have historically shaped biodiversity in regions.
Comparative Biogeographic Studies: The unique historical processes in different temperate regions lead to varying levels of biodiversity despite similar areas.
Conclusion
A comprehensive overview focuses on island biogeography, latitudinal species richness patterns, and historical influences affecting biodiversity, including practical applications in geographic information systems (GIS).