3.1.8.1 = THE GENETIC CODE DNA IS UNIVERSAL, NON-OVERLAPPING & DEGENERATE: nature of genetic code
3.1.8.1 = THE GENETIC CODE
DNA IS UNIVERSAL, NON-OVERLAPPING & DEGENERATE: nature of genetic code
Genetic code: the sequence of DNA triplets bases (or mRNA codons) that code for a specific amino acid.
triplet refers to DNA whereas a codon refers to RNA
The nature of genetic code:
Triple code / 3 bases to each code (each triplet of bases codes for one amino acid)
DNA triplet: sequence of 3 bases coding for specific amino acid, E.g. UAU codes for tyrosine
Universal: The same specific DNA base triplets codes for the same amino acids in all living organisms e.g. UAU codes for tyrosine in all organisms
Non-overlapping: each DNA triplet or mRNA codon contains three bases that codes for a unique amino acid. DNA is discrete; each base from any triplet / codon gene is separate from each other and can only be used once and in only one triplet
Degenerate: The same amino acid can be coded for by more than one base triplet, e.g. tyrosine can be coded for by UAU or UAC
Codon: sequence of three bases (triplet)
Protein synthesis:
Transcription (like a scribe): DNA (nucleotides) → mRNA (nucleotides)
Translation (translating from one language to another): mRNA (nucleotides) → amino acids / protein / polypeptides (amino acids)
3.1.8.2 = POLYPEPTIDE SYNTHESIS
Protein synthesis overview: 2 stages
Transcription; Production of mRNA from DNA
Nucleus
Translation; Production of polypeptides form the sequence of codons carried by mRNA
Cytoplasm on ribosomes
Messenger RNA (mRNA)
Made by transcription in the nucleus
Acts as a template for translation in the cytoplasm
Sequence of bases on RNA determines sequence of amino acids in polypeptide chain
Straight chain molecule
Sequence of bases on RNA determined by sequence of bases on DNA
Triplet code = codon
Chemically unstable
So breaks down after a few days
Transfer RNA (tRNA)
Involved in translation
Carries an amino acid
Amino acid binding site
Anticodon = 3 bases
Anticodon bases complementary to mRNA codon
Each tRNA specific to one amino acid, specific to its anticodon
Single polynucleotide strand
Folded - 3 hairpin loops = three-leafed clover shape
Hydrogen between specific base pairs holds molecule in this shape
Similarities / difference between structure of mRNA and tRNA molecules
Similarities
Both single polynucleotide strand
Differences
mRNA single helix / straight, whereas tRNA folded into clover shape
mRNA is longer, variable length, whereas tRNA is shorter
mRNA contains no paired bases or hydrogen bonds, whereas tRNA has some paired bases and hydrogen bonds
Role of ATP, tRNA and ribosomes in translation
ATP
Hydrolysis of ATP, to ADP + Pi, releases energy
For the bond between the amino acid and its corresponding tRNA molecule
Amino acid attaches at amino acid binding site
For peptide bond formation between amino acids
tRNA
tRNA attaches to and transports a specific amino acid, in relation to its anticodon
tRNA anticodon complementary base pairs to mRNA codon, forming hydrogen bonds
Two tRNAs bring amino acids together for the formation of a peptide bonds
About 60 types of tRNAS to carry 20 different amino acids
Genetic code is degenerate
Ribosomes
Attaches to mRNA and houses tRNA, allowing codon-anticodon complementary base pairing
Allows peptide bonds to form between amino acids
Relating the base sequence of nucleic acids to the amino acid sequence of polypeptides, when provided with suitable data about the genetic code
tRNA anticodons are complementary to mRNA codon
Eg mRNA codon = ACG; tRNA anticodon = UGC
mRNA sequence of bases / codons are complementary to sequence of bases / triplets on DNA template strand
Eg mRNA base sequence = ACG UAG AAC; DNA base sequence = TGC ATC TTG
In RNA, uracil replaces thymine
You may then have to relate this to amino acid sequences
Interpreting data from experimental work investigating the role of nucleic acid in protein synthesis
Protein synthesis: transcription
In nucleus
DNA double helix unzipped by DNA Helicase
Breaking the hydrogen bonds
Forming 2 template strands
Free RNA nucleotides attach to the template strand’s exposed nucleotides
By complementary base pairing (U replaces T in RNA)
H-bonds reform
RNA polymerase joins adjacent mRNA nucleotides together in a condensation reaction forming phosphodiester bonds to form premRNA
When RNA polymerase reaches stop codon, mRNA (prokaryotes) and pre-mRNA (eukaryotes) detaches from DNA
Post transcriptional modification:
Eukaryotic genes = pre-mRNA (messenger) containing exons and introns.
Exons = coding regions
Introns = non-coding regions
pre-mRNA goes into splicing
Introns removed
Exons spliced together in different combos for different proteins and exported
Leaving just the exons and forming mRNA
mRNA now leaves nucleus via nuclear pore for translation in cytoplasm
Prokaryotes vs Eukaryotes
Prokaryotes → transcription :DNA → mRNA
No introns
No splicing → mRNA produced directly from DNA
Eukaryotes: DNA → premRNA → mRNA
Introns are removed (splicing) to form mRNA
Protein synthesis - translation
Codon: triplet of bases on mRNA that codes for a specific amino acid
Anticodon: triplet of bases on tRNA that is complementary to codon on mRA
Sequence of mRNA codons determines sequence of amino acids
tRNAS carry specific amino acids, in relation to their anticodon
At the ribosome, tRNA codon binds to mRNA codon at a start codon
tRNA anticodon attach to mRNA codon by complimentary base pairing
Hydrogen bonds formed
First codon = start codon
Ribosome holds tRNA in place
Two amino acids joined by condensation, forming a peptide bond
Using energy from ATP
tRNA detaches (without its amino acid), ribosome moves along mRNA to next codon
Continues until stop codon (polypeptide released)
ATP has two roles in translation. It is required to provide energy to attach amino acids to tRNA and also to attach amino acids together.
3.1.8.3 = PROTEIN FOLDING
Protein folding: process by which a polypeptide folds into its characteristic three-dimensional structure.
Protein folding is determined by the amino acid sequence of the polypeptide. Specialised proteins, called
chaperones, assist in the folding of other proteins.
Students should be able to:
• relate base sequences of nucleic acids to the amino acid sequence of a polypeptide when provided with suitable information relating to the genetic code.
Students will not be required to recall in written papers specific codons and the amino acids for which they code.
Primary structure of polypeptide is coiled / folded to produce secondary structure
Interactions between R-groups further coil / fold polypeptide to produce tertiary structure (3D shape)
Different polypeptide chains + non-protein groups link to form functional proteins
This is a quaternary structure
*** Some amino acids have neutral non-polar R groups that are hydrophobic, hydrophobic parts of a molecule interact with each other
The sequence of amino acids position of the R groups in the polypeptide determines how a protein is folded.
Disulphide bonds (eg in cystine with another cystine)
Acidic / basic R groups form ionic bonds
Hydrophobic R groups aggregate together / join other hydrophobic molecules → protein is mis-folded → non functional → build up of misfolded protein can cause disease eg alzheimer's disease
Chaperone proteins: ensure polypeptide chains synthesised on ribosomes fold correctly. Some found in endoplasmic reticulum → facilitate folding and assemlby of membrane and sexeerory proteins
E.g. HSP 70. Bind to hydrophobic regions on polypeptides as they are being formed and prevent incorrect hydrophobic interactions occurring before polypeptide chain is completed
E.g. HSP 60: chaperonins, large cylindrical proteins with central compartment. Polypeptide chain fits into central compartment and isolated from ogre proteins molecules → prevented from interacting with them. Interaction between chaperonin and polypeptide enable the protein to fold correctly