loss of biodiversity
what is causing biodiversity variation and change?
natural processes
disturbance
competition
predation
these usually maintain the biodiversity
human caused drivers:
land cover change
chemical release/pollution
overfishing/overharvesting
climate change
etc. (other human impacts)
global cumulative human impact (CHI)
87-90% of the ocean surface has been impacted by humans
Halpern et al. 2019 data show that some areas have
hgh CHI + fast increasing change
high CHI + decreasing change
low CHI + decreasing change
human impacts are global, and the pace of change differs among regions
marine defaunation
marine defaunation started tens of thousands of years later than on land
defaunation is less severe in the ocean than on land
humans have impacted 87-90% of ocean surface
marine fish abundance has decreassed by 38% since 1970
marine habitats such as seagrass and mangroves have decline by > 2/3
land-cover change
human casued land cover change typically decreases species richness
but by creating a more heterogenous habitat, richness can increase
during recovery after disturbance, richness can peak at intermediate stages, then decline at the climax stage
direction of impact depends on disturbance stage and habitat structure
chemical release/pollution
harmful pollutants (mine tailings, insecticides, acid rain) usually decrease richness
fertilizer effects are more complex
richness can increase or decrease
biomass of producers and detritivores usually increases
overharvesting
nonselective harvesting → drastically decreases biomass and species richness
selective harvesting (e.g. targeting top predators) can cause predator release → sometimes increasing biomass or richness
impact direction varies
climate change
climate change alters physical and chemical properties of oceans.
species responses vary within and between groups
ocean warming leads to:
increases in warm water species
poleward shifts
depth shifts
responses where geographical barriers block movement
other impacts listed:
changes in reproduction, growth, survival
changed in ocean currents affecting dispersal
calanus ratio in the north sea has increased, illustrating warm-water dominance with warming
ocean acidification
OA leads to declines in calcification in corals and coccolithophores
it affects growth, reproduction, and organisms with calcium carbonate shells/skeletons
some positive effects exist, e.g., algae (enhanced photosynthesis or fewer predators)
many species are experimentally sensitive to OA
little filed evidence so far; warming dominates current patterns
some field studies (calcifying plankton, foraminifera) show patterns consistent with OA
undersaturation is occuring faster than expected in the western arctic
projections suggest more impacts in the future
the source of extinction threats
its mentioned from the given papers which we will come to soon
global policy frameworks
a new global framework (CBD draft 2030)
target includes:
30% protection of land and sea
50% reduction in invasive species introduction
reduce nutrients by half, pesticides by two thirds
eliminate platic waste discharge
UN SDGs relevant
goal 13: climate action
goal 14: life below water
goal 15: life on land
debate on biodiversity change
globally: extinctions increasing
locally: many argue species richness is not declining
biodiversity trends are complex and vary with scale
important to ask: what is diversity
types of biodiversity
species diversity → many species with similar abundances
genetic diversity → how closely related individuals are; needed for adaptation
ecosystem diversity → number of ecosystems in a region
functional diversity → range of behaviours / ecological function of species
biodiveristy metrics: ⍺- and β diversity
⍺-diversity
number of species at a site
species richness
most commonly measured
β-diveristy
change in species composition over space/time
turnover
biodiversity trends at different scales
global scale:
speciation ≠ extinction balance → extinctions increasing
⍺-diversity decreasing
biogeographical scale
alpha-B (biogeographic scale alpha diversity) increasing
regional (meta-community) scale
trends hypothesized, not well measured
local scale
no significant trend in alpha diversity across studies
large variation among sites
trends in spatial β-diversity
β-diveristy tends to decrease → sites become more similar over time
this is homogenization
leads toward a more spatially homogenous planet
winners (generalists) vs losers (specialists)
generalists adapt better to changes → tey increase
specialists decline → they lose
this lead to functional homogenisation
conclusions from the lecture
biodiveristy trends depend on scale
multiple metrics must be used
be critical - biodiversity numbers can be misinterpreted
trend: moving towards a more homogenous planet
why biodiversity matter?
ecosystem functioning depends on:
species richness
species composition
functional group richness
genetic diversity
ecosystem stability depends strongly on species richness/composition
therefore:
biodiversity declines → less stable ecosystems
global extinction rates
90% of marine organisms extinct over earth history
background rate = 0.01-2 extinctions per million species per yeat
current = 100x higher
humans dominate biogeochemical cyces and biodiversity patterns
ocean acidification has been associated with three of five past mass extinctions
first time such changes are human driven
state of the ocean (Luypaert et al. summary slide)
humans impacted 87-90% of oceans
marine fish abundance down 38% since 1970
coastal habitats (seagrass, mangroves) depleted by two thirds
CO2 up 40% vs pre industrialisation
degraded ecosystems → lost ecosystem services
increased reliance on ocean as land degrades → more pressure
extinction of marine species (Webb & Mindel summary slide)
marine extinctions appear lower than terrestrial
but nuber of marine species is uncertain (estiamtes range: 300,000 - 2.2 million)
only 240,000 marine species described → 11-78% known
extinction risk assesment depends on taxonomic knowledge
IUCN focuses on well-known groups
apparent low marine extinction rate due to lack of knowledge, not low risk
why non-scientific communities doubt biodiversity decline?
public thinks changes are natural fluctuations
people lack trust in data
need clear communication and evidence to build trust
shifting baseline syndrome
concept: each geenration acceps a degraded state as normal
introduced by pauly (1995)
leads to underestimating long-term losses
examples shown from Oslo fjord:
histroical sprat fisheries
many whales in the fjord in 1890
old fishing maps showing abundant fishing spots
lecture summary:
multiple human dirvers interact to change biodiveristy
impacts can increase or decrease richness depending on mechanism
global patterns show strong defaunation but complex local trends
need to measure alpah and beta diveristy across scales
the planet is becoming more homogenous
generalists rise, specialists decline
biodiveristy underpins ecosystem functino and stability
marine extinction appears low but is actuaally underestimated
shifitng baselines distort human perception of loss
the papers - Luypaer et al. 2020, Webb and Mindel 2015, Herbert-Read et al. 2022
integrated explanation
together these papers provide a full picture of where marine biodiveristy stands today, how threatenened it actually is, and what fufutre pressures are coming
the state of marine biodiversity today
human impact is nearly universal
87-90% of the global ocean surface is impacted by human activities
these include exploitation, habitat degredation, climate change, and pollution
major delcines already documented
marine fish abundance has declined by 38% compared to 1970
coastal habitats such as seagrass meadows and mangroves have been depleted by more than two-thirds
atmspheric CO2 has increased by 40%. relative to pre industrial levels, accelrating ocean acidification
possible mass extinction trajectory
over 90% of marine organisms that have ever lived are extinct (natural background extinction)
but current extinction rates are 10x the bacjground rate, suggesting we may be entering a human-driven mass extinction
ocean services are weakening
as ecosystems degrade, the oceans’s ability to deliver ecosystem services decline
if land based ecosystems degrade further, pressure on marine resources will increase even more
marine biodiversity is declining significantly, but the full extnet is hard to measure due to knowledge gaps
why marine extinction appear lower than on land
marine species appear to have lower extinction and extinction-risk levels - but this is misleading
apparent extinction rates
only 19-24 marine species out of > 850 documented global extinctions → this looks like a low marine extinction rate
but the paper shows that this result is biased by limited knowledge
most marine species are not described
estimated number of marine species ranges from 300,000 to 2.2 million
only about 240,000 have been formally described → this means 11-78% of marine species remain unknown
since extinction risk can only be assessed for described species: we underestimate marine extinction risk simply because we lack data
low assessment effort
only 3% of described marine species have been assessed by the IUCN
many marine taxa have no assessments at all
when we do have data, risk is similar to land
when the paper compares well-studied groups (both marine and terrestrial: 20-25% fo species are threateneed with extinction in both realms
in other words: the true marine extinction risk is simialr to terrestrial levels but hidden due to low taxonomic and conservation assessment efforts
data-deficient species
28.6% of marine IUCN assessments are data deficient
DD species often have traits associated with high extinction risk
the ocean doesnt have fewer threateneed species - we just know far less about them. The low risk narrative is an illusion created by lack of data
emerging issues that will impact biodiversity in the next 5-10 years
the horizon scan identifies 15 emerging issues likely to storngly affect marine biodiverisry soon. These issues fall into three categories
ecosystem impacts:
wildfire impacts on coastal/marine ecosystems
fires release nutrients, metals and particles that wash into the ocean
can cause eutrophication, mortality of benthic invertebrates, or temporary productivity increases
coastal darkening:
increased turbidity and dissolved organic matter reduce light penetration
affects photosynthesis, species composition, and coastal habitat function
increased metal toxicity due to ocean acidification
lower pH increases bioavailability and toxicity of chemicals
affects marine organisms and potnetially human health (via seafood)
declining equatorial biodiversity
equatorial regions are losing species as thermal zones shift poleward
creates a dip in diversity around the equator
lower nutritional content of fish due to climate driven changes
changes in phytoplankton fatty acids affect foodwebs and human nutrition
resource exploitation:
marine collagens
increasing demand for collagen may drive harvesting og sponges, jellyfish, sharks, etc.
risk of increased pressure on non-target species
expanding trade in fish swim bladders
extremely lucrative trade drives overexploitation
bycatch threatens species such as sharks, rays, turtles
fishing of mesopelagic species
huge biomas but critical for the biological carbon pump
large scale harvesting could disrupt carbon sequestration
deep sea lithium extraction
higher lithium demand (e.g. batteries) may shift mining to deep sea brines
deep brine ecosystems contain many endemic and largely undescribed species
new technologies
colocation of marine activities
combined uses (wind farms + aquaculture) require new regulatory frameworks
floating marine cities
could alter species movement, increase invaive species, and raise governance issues
trace element contamination form green technologies
battery waste and production release nickel, cobalt, and other toxic elements
new underwater tracking systems
could improve study of non surfacing animals, but ecological impacts are unknown
soft robotics
useful for deep sea sampling but may introduce pollutants or disturb organisms
biodegradable materials
green plastics may still causes harm; theri long term impact is unknown
new threats are emerging rapidly, mnay of which are poorly understood or not yet included in management plans
take home message
marine biodiversity is already declining severly, is more threateneed than it appears due to huge knowledge gaps, and will face additional emerging threats in the coming years
together the three papers present a consistent picture:
the ocean is heavily impacted by real extinction risk is underestimated
new, poorly understood pressures are emerging rapidly
this reinforces teh lectures themes:
biodiversity is declining globally
trends differ across spatial scales
generalists may increase, specialists decline
management decisions depend on good data - but marine biodiversity is understudied
future pressures may increase homogenization and degredation