Overview of Animal Diversity
Overview of Animal Diversity
What are Animals?
Definition of Animals:
Multicellular
Heterotrophic
Eukaryotic
Develop from embryonic layers
Exceptions to Living Things:
Example: Plasmodium falciparum
Considered by many to be an animal despite being unicellular
Previously covered with protists
Importance of Criteria:
Most animals fit the established criteria.
Nutritional Modes
How Animals Eat:
Animals cannot synthesize their own organic materials (unlike plants).
Plants: autotrophic
Fungi: obtain nutrition by decomposing organic matter
Most animals: dissolve materials within their bodies for nutrition.
Cell Structure and Specialization
General Cell Structure:
Most animals are eukaryotes and multicellular.
Lack of Cell Walls:
Unlike plants and fungi, animals do not have cell walls.
Animals use proteins, including collagen, in their cell membranes to connect cells.
Tissue Specialization:
Presence of specialized tissues (e.g., nervous and muscle tissues).
Unique capabilities that fungi and plants do not possess.
Reproduction and Development
Modes of Reproduction:
Most animals reproduce sexually, with many exceptions.
Dominance of Diploid Stage:
Diploid stage is the prevalent state in the animal lifecycle.
Sperm and egg production occurs via meiosis.
Zygote Development:
Zygotes are formed when small, flagellated sperm fertilizes larger, nonmotile eggs.
Zygotes undergo mitotic divisions:
Cleavage: series of mitotic divisions sans cell growth.
Blastula Formation: A multicellular stage that forms a hollow ball of cells, leading to gastrulation.
Gastrulation Explained:
The process where one end of the embryo folds inward.
Forms multiple layers of embryonic tissues:
Ectoderm: outer layer.
Endoderm: inner layer.
Early Embryonic Development in Animals
Stages in Embryonic Development:
Zygote undergoes cleavage to form a multicellular blastula.
Blastula develops into a gastrula, which involves an indentation (blastopore).
Larval Stages and Metamorphosis
Adult Development:
Many animals develop directly into adults but others have larval stages.
Definition of Larva:
A sexually immature form, distinct from the adult.
Engages in different feeding behaviors from adults.
Metamorphosis:
Developmental transformation from larva to adult.
Some animals undergo a single larval stage, while others may have multiple.
Examples of Larval Development
Dragonfly Life Cycle:
Stages include egg, naiad (larva), and adult.
Animal Diversity
Estimates of Animal Species:
Approximately 1.3 million extant (living) species, with many more undiscovered.
Tremendous Diversity of Animal Types:
Examples include corals, cockroaches, crocodiles, humans, and snails.
Common Ancestor of Animals:
Likely lived around 770 million years ago.
Closest related protist group: Choanoflagellates.
Ediacaran Biota:
Members resemble various animal forms including mollusks, sponges, and cnidarians.
Paleozoic Era
Timeline: 541-252 million years ago.
Cambrian Explosion:
Major increase in animal diversity occurring between 535-525 million years ago.
Emergence of bilaterans.
Definition of Bilaterans:
Animals exhibiting bilateral symmetry.
Notable exceptions: sponges and cnidarians.
Post-Cambrian Period
Diversity Growth:
Continued increase in animal diversity during the Ordovician, Silurian, and Devonian periods.
Existence of mass extinctions.
Marine Dominance:
Vertebrates were the dominant sea animals.
Fish and arthropods (e.g., spiders, millipedes, centipedes) were the first to colonize land.
Mesozoic Era
Timeline: 252-66 million years ago.
Evolutionary Developments:
Further diversification as new habitats were occupied.
Creation of coral reefs.
Reptiles returned to aquatic environments (e.g., Plesiosaurs).
Evolution of wings among tetrapods (birds and pterosaurs).
Presence of dinosaurs (both herbivorous and predatory).
Rapid diversification of mammals and angiosperms.
Cenozoic Era
Timeline: 66 million years ago to present.
Climate and Ecological Changes:
Extinction of non-flying dinosaurs and marine reptiles.
Decline in climate temperature, leading to different vegetation types.
Replacement of dense forests in Africa with savannas and open woodlands which benefited primates, leading to the evolution of human ancestors.
Body Plans and Symmetry
Definition of Body Plan:
Morphological and developmental traits integrated into a functional whole.
Misconception of “plan”:
Animals did not consciously design their morphology (e.g., blobfish, naked mole rat).
Types of Symmetry
Symmetry Variations:
No Symmetry: Example: sponges.
Radial Symmetry: Predominantly in cnidarians (e.g., jellyfish, sea anemones).
Bilateral Symmetry: Most animals characterized by two axes.
Dorsal (top) and ventral (bottom).
Anterior (front) and posterior (back).
Lifestyle Associations:
Radially symmetrical animals tend to be sessile.
Bilateral animals are usually more active.
Tissues in Animals
Definition of Tissues:
Specialized groups of cells that work together.
Example: sponges lack true tissues; other animals develop tissues during gastrulation.
Germ Layers Defined:
Ectoderm: outer layer of the embryo.
Endoderm: innermost layer giving rise to organ linings.
Diploblastic: Radially symmetrical animals possess 2 layers (ectoderm and endoderm).
Triploblastic: Bilateral animals with three layers (ectoderm, endoderm, and mesoderm).
Body Cavities
Body Cavities in Triploblastic Animals:
Cavities are located between the digestive tract and body wall, called coelom.
Functionality of Body Cavity:
Provides cushioning for suspended organs and contains fluid to shape the body.
Facilitates organ growth and movement without disruption.
Protostome vs. Deuterostome Development
Key Differences:
Cleavage type, coelom formation, and blastophore configuration differ.
Blastopore: The indentation leading to gastrulation.
Developmental Patterns:
Protostomes: Develop through spiral cleavage, with determinate cleavage meaning early cell fate.
Deuterostomes: Develop through radial cleavage and possess indeterminate cleavage allowing early cells to maintain totipotency (example: identical twins).
Coelom Formation
Archenteron: The gut-forming blind pouch produced during gastrulation.
Coelom Development:
In protostomes: mesoderm solid masses form the coelom.
In deuterostomes: mesoderm buds from the archenteron to form the coelom.
Forming Openings:
In protostomes, the mouth develops first; in deuterostomes, the anus forms first.
Diversification of Animals
Common Characteristics Shared by Animals:
Share a common ancestor.
Sponges as the sister group to all other animals, recognized as the most basal animals.
Eumetazoa: Clade of animals with specialized tissues (e.g., nerve and muscle tissue).
Majority of animals exhibit bilateral symmetry, significant diversification occurring during the Cambrian explosion.
Major clades of bilaterans:
Deuterostomia: includes hemichordates, echinoderms, and chordates.
Lophotrochozoa: includes animals like molluscs with a lophophore (crown of ciliated tentacles).
Ecdysozoa: includes nematodes and arthropods, known for shedding their exoskeletons (ecdysis).