CHapter 11 Flashcards

Chapter 11: Biological Membranes and Transport

Page 1: Introduction

  • Overview of biological membranes and their importance in transport mechanisms.

Page 2: Lipids Aggregation

  • Three major structures formed by lipids:

    • Micelles

    • Liposomes

    • Bilayers

  • Structure formation depends on:

    • Type of lipid

    • Concentration

Page 3: Micelles

  • Description:

    • Formed by amphipathic molecules with larger polar heads than tails.

    • Contain a few dozen to a few thousand lipid molecules.

  • Critical micelle concentration (CMC) is key for micelle formation.

    • Examples: Fatty acids and sodium dodecyl sulfate.

Page 4: Vesicles (Liposomes)

  • Synthetic vesicle membranes can be created in vitro with inserted proteins.

  • Central aqueous cavity can contain dissolved molecules (e.g., drugs).

  • These are useful for transporting molecules and can fuse readily with cell membranes.

  • Small bilayers spontaneously seal into spherical vesicles based on concentration.

Page 5: Membrane Bilayer

  • Forms in aqueous solutions with lipids having polar head groups and more than one tail.

  • Composed of:

    • Two leaflets of lipid monolayers (e.g., phospholipids, sphingolipids).

  • Depicts hydrophobic (tails) and hydrophilic (heads) interactions.

Page 6: Exam Question Example

  • Question: Which change increases micelle diameter?

    • A: Increasing concentration of cholesterol

    • B: Increasing length of acyl chains

    • C: Increasing concentration of polyunsaturated fatty acids

    • D: Using lipids with more polar head groups.

Page 7: Membrane Definition

  • Membranes are complex lipid-based structures that form pliable sheets.

  • Composed of various lipids and proteins, defining boundaries of cells and compartments in eukaryotic cells.

Page 8: Membrane Functions

  • Define boundaries for cells.

  • Import/export nutrients (e.g., lactose) and waste (e.g., toxins).

  • Retain metabolites and ions, sense external signals, provide compartmentalization for reactions.

  • Important for energy storage (proton gradients) and nerve signal transmission.

Page 9: Common Features of Membranes

  • Sheet-like, flexible structure (30–100 Å thick).

  • Typically, two leaflets of lipids (bilayer).

  • Form spontaneously and are stabilized by noncovalent forces, particularly the hydrophobic effect.

  • Asymmetric distribution of lipids and carbohydrates attached outside.

  • Fluid structures where movement is prevalent.

Page 10: Fluid Mosaic Model

  • Proposed by Singer and Nicholson in 1972.

  • Describes the arrangement and movement of proteins within the lipid bilayer, both embedded and loosely associated.

Page 11: Variability in Composition

  • Membrane composition varies across organisms, tissues, and organelles:

    • Abundance and type of lipids and proteins, e.g., absence of cholesterol in mitochondria and abundant galactolipids in chloroplasts.

Page 12: Asymmetry in Membranes

  • Different lipid compositions in leaflets:

    • Outer leaflet often positively charged.

    • Specific functions associated with lipid positioning (e.g., blood clotting, cell signaling).

Page 13: Membrane Structure Highlights

  • Plasma membrane components and associated structures:

    • Golgi apparatus, endosomes, and lysosomes with specific lipid compositions.

Page 14: Exam Question

  • Question: Identify the false statement regarding biological membranes.

    • A: All membranes have the same composition.

    • B: All contain lipids.

    • C: Lipid bilayers are impermeable to polar solutes.

    • D: Membranes typically contain proteins.

Page 15: Membrane Proteins Overview

  • Membrane proteins categorized into:

    • Integral: span the entire membrane

    • Peripheral: loosely attached, interact with aqueous domains.

Page 16: Peripheral Membrane Proteins

  • They associate with polar head groups through ionic interactions.

  • Removed by disrupting interactions (e.g., high salt conditions).

Page 17: Lipid-Linked Proteins

  • Proteins covalently linked to lipids, influencing targeting and membrane function.

  • Examples include glycosylated phosphatidylinositol (GPI) anchors.

Page 18: Integral Membrane Proteins

  • Span entire membrane with hydrophobic transmembrane segments, interact with lipid bilayer.

  • Examples include channels for nutrient transport.

Page 19: Membrane Protein Composition

  • Composition includes various secondary structures, e.g., alpha-helices and beta-sheets.

  • Responsible for various membrane functions.

Page 20: Membrane Protein Functions

  • Functions of membrane proteins:

    • Receptors detecting external signals (e.g., hormones).

    • Ion channels for transporting ions and nutrients.

    • Enzymatic role in lipid metabolism (e.g., ATP synthase).

Page 21: Exam Questions on Membrane Proteins

  • Topics on amphitropic proteins:

    • Functions and characteristics within membranes.

Page 22: Effects on Membrane Fluidity

  • Membrane fluidity influenced by lipid composition and temperature, affecting biological functions.

Page 23: Membrane Dynamics

  • Lateral diffusion of lipids occurs rapidly.

  • Experimental techniques (e.g., FRAP) monitor lipid movement and dynamics.

Page 24: FRAP Methodology

  • Steps involved in the fluorescence recovery after photobleaching for studying membrane dynamics.

Page 25: Special Enzymes and Proteins

  • Flippases facilitate lipid movement across bilayers; significant for membrane asymmetry.

Page 26: Membrane Rafts

  • Lipid distribution is not random; specialized areas with particular lipid and protein clustering.

Page 27: Membrane Fusion

  • Membranes can fuse without exposure to solvent, a critical process in cellular communication.

Page 28: Membrane Transport Mechanisms

  • Transport must be energetically favorable, considering concentration and electrochemical gradients.

Page 29: Types of Membrane Transport

  • Different mechanisms:

    • Simple diffusion for nonpolar substances.

    • Passive diffusion versus facilitated diffusion via transport proteins.

    • Active transport mechanisms driven by ATP.

Page 30: Membrane Properties Summary

  • Membrane fluidity and dynamics are crucial for physiological functions, with varied effects based on lipid structures.

Page 31: Exam Questions on Transport

  • Questions about membrane depolarization, protein embedment, and hydrophobic segments in complex processes.

  • Feedback mechanisms that drive homeostasis and communication at cellular levels.