Male Reproductive System
Male Reproductive System Notes
Objectives
- Understand hormones secreted by Leydig and Sertoli cells in the testes.
- Learn about spermatogenesis.
- Explore mechanisms of erection and ejaculation.
- Study the structure, biosynthesis, transport, metabolism, and actions of testosterone.
- Discuss the regulation of testosterone secretion.
Anatomy of the Male Reproductive System
- Components include:
- Urinary bladder
- Seminal vesicle
- Prostate gland
- Testes and epididymis
- Vas deferens
- Urethra
- Penis
- Scrotum
- Each component has specific functions that contribute to male reproductive health.
Physiologic Anatomy of the Testes
- Testes contain convoluted seminiferous tubules and interstitial Leydig cells.
- Convoluted seminiferous tubules: Site of spermatogenesis where sperm are formed.
- Leydig cells: Secrete testosterone and contain lipid granules.
- The anatomy allows for countercurrent exchange of heat and testosterone, essential for testicular function.
Blood-Testis Barrier
- Sertoli cells and germ cells maintain survival of spermatogenic cells through:
- Tight junctions creating a blood-testis barrier, restricting large molecule passage.
- Facilitates a specific composition of tubular fluid necessary for sperm development.
- Actions of the blood-testis barrier include:
- Protecting germ cells from pathogens.
- Preventing autoimmune responses against germ cells.
Spermatogenesis
- Starts with spermatogonia maturing into primary spermatocytes.
- Primary spermatocytes undergo meiosis to form secondary spermatocytes, then spermatids which develop into spermatozoa.
- Total transformation takes approximately 74 days.
- The process ensures synchronization through cytoplasmic bridges among developing cells.
Structure of Spermatozoa
- Composed of:
- Head: Contains chromosomal material covered by acrosome rich in fertilization enzymes.
- Midpiece: Contains mitochondria providing energy for motility.
- Tail: Facilitates movement toward the ovum.
- Germinal angiotensin-converting enzyme (gACE) may play roles in fertility.
Role of Sertoli Cells
- Support spermatogenesis by:
- Providing androgen binding protein (ABP) and inhibin.
- Converting androgens to estrogens via aromatase.
- Regulate fluid composition in the testes to support sperm maturation.
Hormonal Regulation
- Testosterone and FSH are crucial for testicular function:
- LH stimulates Leydig cells for testosterone production.
- FSH maintains spermatogenesis via Sertoli cells.
- Inhibin feeds back to inhibit FSH secretion, helping regulate sperm production.
Temperature Effects
- Ideal testicular temperature is about 32°C for optimal spermatogenesis.
- Increased temperature (e.g., hot baths, insulation) can result in reduced sperm counts.
Semen Composition
- Avg volume per ejaculate: 2.5–3.5 mL containing about 100 million sperm/mL.
- Semen includes secretions from seminal vesicles, prostate, and bulbo-urethral glands.
- Prostaglandins present but functions are unclear.
Erection Mechanisms
- Initiated by dilation of arterioles and filling of erectile tissue with blood.
- Affected by the release of acetylcholine and nitric oxide (NO) which promote vasodilation via cGMP.
- NO synthase inhibitors can prevent erections, while drugs like sildenafil promote prolonged erections by inhibiting breakdown of cGMP.
Ejaculation Process
- Involves two parts: emission (movement of semen into urethra) and ejaculation (propulsion of semen out).
- Fluid propulsion occurs due to contraction of the bulbocavernosus muscle.
Testosterone Biosynthesis
- Testosterone is derived from cholesterol in Leydig cells and regulated by LH.
- The synthesis involves various hydroxylation and cleavage processes.
Actions of Testosterone
- Develops and maintains male secondary sex characteristics.
- Influences protein synthesis and promotes growth (anabolic effects).
- Feedback inhibition of LH secretion occurs through direct action on the anterior pituitary.
Inhibins and Activins
- Inhibins inhibit FSH secretion, while activins stimulate it.
- Inhibins formed by Sertoli cells and granulosa cells.
Abnormalities of Testicular Function
- Conditions include: cryptorchidism and androgen-secreting tumors.
- Understanding hormonal levels assist in diagnosing related conditions.