Symbioses other than zooxanthellae

  • prokaryotes likely play an important role in the coral holobiont

  • prokaryotes and algae form important symbioses with sponges

  • vertical transmission of prokaryotic symbionts would imply

    • acquisition from parents

  • mode of sponge symbiont acquisition are likely similar to coral-zooxanthellae

  • some bacteria are detrimental to sponges and coral

  • there are numerous macro-symbioses on reefs

    • pearlfish and sea cucumbers

    • obligate symbiosis of gobies and coral

      • gobies as cleaners

    • cleaner shrimp

    • many pelagic species return to reefs for cleaning

    • fish-anemone symbioses