Evolution, phylogenetics, origin (1)
BIO104 Evolution and the Diversity of Life
Quote by Theodosius Dobzhansky: "Nothing in biology makes sense except in the light of evolution."
What is Evolution?
Evolution explains the origin of biological diversity.
Key questions:
Where does diversity in biology come from?
How do new species form?
When do we classify a species as diverging?
Are species always in competition?
What causes species extinction?
Example: Evolution of the genus Homo (Homo habilis, Homo erectus, Homo neanderthalensis, Homo sapiens)
Definitions and Concepts of Evolution
Evolution: A process resulting in changes in the genetic content of populations over time, detailing inherited trait changes through generations.
Misconceptions: Evolution does not aim towards a specific goal or endpoint.
Types of Evolution
1. Microevolution
Changes at or below species level.
Involves variation in traits and frequencies among population members due to ecological shifts, environmental changes, and interspecies interactions.
May lead to new species over time but often results in fluctuating trait frequencies based on selective pressures.
2. Macroevolution
Changes occurring above the species level.
Macroevolutionary changes typically observed over long time scales and inferred from fossil records, phylogenetic studies, and microevolution patterns.
Helps address present ecological phenomena, such as climate impacts on species survival.
The Theory of Evolution by Natural Selection
Origin of Species by Charles Darwin outlines key evolutionary theories:
Evolution as such: Constant change in the world; organisms evolve over time.
Common descent: All organisms share a common ancestor.
Multiplication of species: Species diversify through splitting and budding.
Gradualism: Evolution occurs gradually, not through sudden changes.
Natural Selection: Adapted traits enhance survival and reproduction across generations.
Evidence for Evolution Theories
Theory 1: Evolution as Such
Fossil evidence demonstrates the non-constant nature of organisms.
Theory 2: Common Descent
All organisms trace back to a single origin:
Example: Evolution of whales showcases transitional forms over millions of years.
Theory 3: Multiplication of Species
Geographic isolation leads to new species formation, illustrated by different finch species in Hawaii.
Theory 4: Gradualism
Evolutionary change is a slow, consistent process rather than bursts of new forms.
Theory 5: Natural Selection
Variations arise from mutations; survival favoring well-adapted individuals leads to population changes over time.
Key Concepts: Variation, Inheritance, Selection, Time, and Adaptation (VISTA).
Phylogenetics
Importance: Understanding evolutionary relationships, classification, forensics, and conservation efforts.
Phylogenetic Trees: Represents evolutionary relationships.
Properties of a Phylogenetic Tree
Topology: Indicates how organisms are related.
Branches: Represent genetic change; longer branches indicate more divergence.
Nodes: Points of evolutionary history.
Root: Most recent common ancestor of taxa, demonstrating evolutionary direction.
Outgroup rooting: Utilizes distantly related sequences for accurate tree rooting.
Understanding Life and Evolutionary Origins
Chemical Origins of Life
Proposed theories include:
Panspermia: Life or its building blocks came from space.
Spontaneous Generation: Disproven idea that life arose from non-living matter.
Chemical Evolution: Suggests that life originated from simple chemical processes leading to complexity.
Urey-Miller Experiment: Demonstrated that organic compounds can form under prebiotic conditions.
Criteria of Life
Characteristics include cell structure, metabolism, and reproduction capabilities.
LUCA (Last Universal Common Ancestor)
Hypothetical ancestor linking abiotic Earth and early life forms, thought to be a complex community rather than a simple organism.
Predicted properties: unicellular, RNA as genetic material, adaptability to extreme environments.
Cell Theory
Developed by Theodor Schwann and Matthias Jakob Schleiden:
All organisms are made of cells.
Cells are the structural and functional units of life.
Cells arise from pre-existing cells.
Modern interpretation includes that cellular activity depends on independent cell activities.
Prokaryotic vs. Eukaryotic Cells
Prokaryotic Cells
Characteristics: Smaller in size, unicellular, no nucleus, circular DNA, lack membrane-bound organelles.
Domains: Bacteria and Archaea.
Eukaryotic Cells
Characteristics: Larger, more complex, nuclei present, linear DNA, membrane-bound organelles.
Examples include plant and animal cells.
Bacterial Structures
Key structures: capsule, cell wall, plasma membrane.
Cell Wall Variability:
Gram-positive vs. Gram-negative bacteria, different structure and staining properties.
Archaea Properties
Distinct from bacteria, live in extreme environments, possess unique biochemical and genetic traits.
Cell walls composed of pseudopeptidoglycan, can utilize diverse energy sources.