Principles of Pharmacology - Week 6 Seminar Notes

Topic = Central Nervous System part 1

Regions of the brain and CNS organization

  • CNS includes the brain and spinal cord; major regions are the cerebrum, diencephalon, brainstem, cerebellum, and spinal cord.

  • Central nervous system (CNS) is organized into distinct functional units that coordinate sensation, movement, emotion, and cognition.

  • Understanding these regions helps explain pharmacological targets and disease states in pharmacology.

Cerebrum and cerebral cortex

  • Cerebrum: largest, uppermost part of the brain; divided into left and right hemispheres connected by the corpus callosum (a thick bridge of axons).

  • Cerebral cortex: outer grey matter with inner white matter (medulla).

  • Primary roles: perception and interpretation of sensation, initiation of skeletal muscle movement, communication, intellect, abstract thought.

Cerebral cortex – lobes and functional areas

  • Four main lobes:

    • Frontal lobe: muscle movement, motor components of speech, abstract thinking, problem solving.

    • Parietal lobe: sensory modalities (touch, pressure, pain, temperature, vibration).

    • Temporal lobe: hearing, learning, memory, language functions.

    • Occipital lobe: vision.

  • Neural connections between lobes enable integration, communication, and coordination.

Detailed cortical regions (illustrative schematic from Fig. 8.7)

  • Central sulcus separates frontal from parietal lobes; precentral gyrus houses Primary Central Motor Cortex; postcentral gyrus houses Somatosensory Cortex.

  • Premotor cortex, Broca’s area (motor speech), Wernicke’s area (sensory speech) are key functional areas.

  • Taste cortex, auditory cortex, and visual cortex are located in their respective lobes; gyri and sulci increase cortical surface area.

Cerebral medulla and basal ganglia

  • Cerebral medulla: inner white matter composed of myelinated axons; conducts signals between cortex and other CNS regions.

  • Basal ganglia: grey matter structures within white matter; regulate motor activity, initiate and smooth voluntary movement; part of extrapyramidal system (EPS), crucial for gross motor control and muscle tone.

Diencephalon

  • Includes thalamus and hypothalamus; located at the back of the forebrain, below the cerebral hemispheres and above the brainstem.

  • Thalamus: relay for sensory and motor impulses; regulates consciousness, sleep, and alertness.

  • Hypothalamus: regulates appetite, temperature, fluid balance, hormonal control, biological rhythms; integrates autonomic nervous system and endocrine function; closely linked to emotion and survival behaviours.

Brainstem and spinal cord

  • Brainstem comprises midbrain, pons, and medulla oblongata.

    • Midbrain: head of the brainstem; contains several reflex centers.

    • Pons: regulation of respiration and sleep.

    • Medulla oblongata: autonomic control over cardiac, vasomotor, and respiratory functions; also involved in swallowing, coughing, vomiting, gagging.

  • Spinal cord: conduit for sensory and motor nerves between brain and body; major conduction pathway for CNS signals.

Cerebellum

  • Means “little brain”; located behind the brainstem below the cerebrum.

  • Coordinates voluntary movements: posture, balance, coordination, speech.

  • Fine-tunes ongoing movements and provides quality control; involved in planning and decision-making related to movement.

Functional components of the brain

  • Reticular formation: network of nerve fibres extending through brain and spinal cord; connects many brain areas; regulates motor control, cardiovascular control, sleep, wakefulness.

    • Contains inhibitory and excitatory neurons; Excitatory nerves form the Reticular Activating System (RAS).

    • Susceptible to modulation by stimulants (e.g., amphetamines) and depressants (e.g., alcohol, barbiturates).

  • Limbic system: interconnected neural network regulating emotions and behavior; functionally connected rather than strictly anatomically defined.

    • Key structures: Hippocampus (long-term memory formation) and Amygdala (emotional responses such as fear, anger, anxiety, reward/punishment).

    • Dysfunctions linked to addictions (drug abuse, smoking) and negative emotional states; anxiolytics and some antidepressants can stabilize limbic activity.

Major brain chemical transmitter systems

  • Catecholamines: Adrenaline, Noradrenaline, Dopamine.

  • Indolamines: Serotonin, Melatonin.

  • Monoamines: Dopamine, Serotonin, Noradrenaline, Adrenaline (noradrenaline and adrenaline are sometimes grouped with catecholamines).

  • Other key transmitters: GABA (inhibitory), Glutamate (excitatory), Acetylcholine (ACh), Histamine, Endorphins and Enkephalins, Adenosine, Substance P.

  • Transmitter families are often categorized by receptor families and physiological roles.

CNS neurotransmitters – overview and receptor considerations

  • Adrenaline and Noradrenaline (catecholamines)

    • Involved in fight-or-flight, mood regulation, arousal.

    • Adrenergic receptors: α and β subtypes.

  • Acetylcholine (ACh)

    • Pathways controlling voluntary movement, memory, learning, alertness.

    • Cholinergic receptors: muscarinic and nicotinic.

  • Dopamine

    • Predominantly in nigrostriatal pathway (voluntary movement), plus mesolimbic, mesocortical pathways involved in behavior, reward, motivation, addiction, endocrine control via tuberoinfundibular pathway.

    • Receptors: D1-like (D1, D5) and D2-like (D2, D3, D4).

  • Serotonin (5-HT)

    • Wide CNS distribution; regulates mood, arousal, eating, vomiting, cognition, sleep, pain, temperature; can be excitatory or inhibitory depending on receptor subtype.

  • GABA (γ-aminobutyric acid)

    • Brain’s major inhibitory neurotransmitter; widespread control of excitability.

    • Receptors: GABAA (ionotropic) and GABAB (metabotropic).

  • Glutamate

    • Brain’s major excitatory transmitter; receptors include NMDA, AMPA, kainate (ionotropic) and mGluRs (metabotropic); essential for learning, memory, and synaptic plasticity.

  • Histamine

    • Involved in wakefulness, arousal, and immune responses; receptors H1–H4 with variable excitatory/inhibitory roles.

  • Endorphins and Enkephalins (opioid system)

    • Modulate pain and mood via μ, δ, κ opioid receptors; generally inhibitory.

  • Adenosine

    • Generally inhibitory; regulates sleep, promotes relaxation, provides neuroprotection; acts via A1, A2A, A2B, A3 receptors.

  • Substance P

    • Neurokinin receptors (primarily NK1); excitatory; involved in pain transmission.

CNS neurotransmitter receptor and functional relationships (selected highlights)

  • Acetylcholine

    • Receptors: nicotinic (ionotropic) and muscarinic (metabotropic).

    • Roles: muscle contraction, cognitive function, autonomic regulation.

  • Dopamine

    • Receptors: D1-like (D1, D5) and D2-like (D2, D3, D4).

    • Generally excitatory in CNS; controls mood, reward, motor function, and cognition.

  • Serotonin (5-HT)

    • Multiple subtypes (5-HT1 to 5-HT7); can be excitatory or inhibitory depending on receptor.

  • Noradrenaline/Adrenaline

    • Adrenergic receptors: α and β; generally excitatory in CNS; involved in arousal and mood.

  • Glutamate

    • Receptors: NMDA, AMPA, kainate (ionotropic); mGluRs (metabotropic).

    • Functions: learning, memory, synaptic plasticity; excitatory transmission.

  • GABA

    • GABAA and GABAB receptors; inhibitory control of neuronal excitability.

Brain regions and transmitter associations (Table-style synthesis)

  • Acetylcholine: Cognition, skeletal muscle movement, memory; associated regions include cerebral cortex, thalamocortical tracts, pyramidal pathways, and the reticular activating system; linked to disorders like Parkinson's disease and dementia when dysregulated.

  • Dopamine: Behaviors, reward pathways, motor control; associated with extrapyramidal pathways and limbic system; involved in emesis and hormone release via hypothalamus.

  • Noradrenaline: Arousal, wakefulness, mood, appetite, hormone release; linked to reticular activating system and hypothalamus.

  • Serotonin: Arousal, sleep, mood, appetite, temperature; widespread in CNS; related to depression, eating disorders, insomnia.

  • GABA: Broad inhibitory control across CNS; abundant in cortex and basal ganglia.

  • Glutamate: Learning, memory, consciousness; widespread excitatory transmission; epilepsy and neurodegenerative processes linked to excitotoxicity.

Disease states of the CNS – general concept

  • CNS disorders often arise from neurotransmitter imbalances:

    • Hyper-excitable neurons (excess excitation) can trigger seizures.

    • Excessive neurotransmitter binding to postsynaptic receptors can contribute to psychoses.

    • Deficits in neurotransmitter levels (e.g., dopamine in Parkinson’s disease) contribute to neurodegenerative processes.

Neurotransmitter balances in CNS disorders (conceptual mappings)

  • Parkinson's disease: Dopamine deficiency with relative acetylcholine excess; motor symptoms prominent.

  • Depression: Monoamine (NA, 5-HT, and possibly DA) dysregulation.

  • Schizophrenia: Dopamine imbalance with hyperactivity in mesolimbic pathways and hypoactivity in mesocortical pathways; dopamine, glutamate and serotonin systems implicated.

  • Dementia: Acetylcholine deficiency and broader neurotransmitter changes.

Sedative and anxiolytic drugs – overview

  • Sleep disorders and anxiety are common targets for CNS depressants and anxiolytics.

  • Short-term hypnotics can improve sleep, but may disrupt normal sleep architecture and carry risks of dependence and tolerance.

  • Anxiety disorders involve GABA, noradrenaline, serotonin, and dopamine systems; limbic structures (amygdala, hippocampus) are key targets.

Sleep disorders and therapeutic approach

  • Sleep is essential; sleep disturbances affect daily life.

  • Initial approach: identify underlying cause, optimize sleep hygiene, review medical/psychiatric history.

  • Hypnotics may provide temporary relief but can disrupt normal sleep cycles.

Anxiety and pharmacotherapy – distinctions

  • Anxiolytics vs hypnotics:

    • Anxiolytics: lower CNS activity to reduce anxiety; lower doses.

    • Hypnotics: promote sleep with higher sedative effects.

  • Sedatives may be used for anxiety or insomnia; the dose and duration differ.

Sedative drugs – pharmacology basics

  • Barbiturates (sedative-hypnotics) and benzodiazepines act to enhance GABAergic inhibition; benzodiazepines mainly act on GABA_A receptors.

  • Zolpidem and Zopiclone are non-benzodiazepine hypnotics that act on GABA_A receptors but with different specificity.

  • Buspirone is an anxiolytic with a less sedative profile; mechanism is not fully clear but distinct from benzodiazepines.

  • Flumazenil reverses benzodiazepine toxicity; short half-life necessitates IV administration.

  • Other hypnotics include Diphenhydramine, Doxylamine (sedating antihistamines) and Melatonin (circadian rhythm regulation).

Benzodiazepines – clinical features and pharmacokinetics

  • Examples: diazepam, alprazolam, temazepam.

  • Mechanism: enhance GABAergic inhibition via GABAA receptors; broad CNS effects due to widespread GABAA presence.

  • Indications: anxiety, insomnia, panic, seizures, anesthesia adjunct, alcohol withdrawal.

  • Adverse effects: dependence, altered sleep architecture, daytime drowsiness, cognitive/memory impairment, increased fall risk in older adults.

  • Pharmacokinetics:

    • Highly lipophilic: rapid, complete absorption.

    • Can accumulate in fat stores, especially in older patients.

    • Large variation in half-lives; long-acting agents may have active metabolites extending action.

    • Individual responses vary; some benzodiazepines have active metabolites contributing to duration.

  • Tolerance and withdrawal: risk of dependence; withdrawal symptoms are a concern when stopping.

Benzodiazepines – practical considerations

  • Different half-lives influence onset/duration, insomnia vs anxiety treatment, risk of dependence, daytime drowsiness.

  • Tolerability and safety profile are influenced by age, comorbidities, and concomitant medications.

Non-benzodiazepine agents for anxiety and sleep disorders

  • Barbiturates: act on GABA_A receptors; higher risk profile and dependence potential.

  • Zolpidem (hypnotic): GABA_A receptor action; risks include tolerance, dependence, withdrawal.

  • Zopiclone (hypnotic): enhances GABA_A activity; elderly patients are more susceptible to adverse effects.

  • Buspirone (anxiolytic): mechanism not fully defined; generally well tolerated; limited sedation and motor impairment.

Reversal and other agents

  • Flumazenil: benzodiazepine antagonist; short half-life; administered IV for acute toxicity reversal.

Other hypnotic drugs

  • Diphenhydramine and Doxylamine: sedating antihistamines with hypnotic effects.

  • Melatonin: hormone involved in circadian regulation; used as a sleep aid.

Alcohol – CNS depressant effects and withdrawal

  • Pharmacological effects: widespread CNS depression; vascular, GI, renal, nutritional effects.

  • Acute effects: CNS depression; chronic use leads to tolerance and dependence; withdrawal can be debilitating.

  • Management concept: avoidance therapy (where a conditioning stimulus is paired with adverse effects to discourage drug use).

  • Disulfiram (antabuse): used in alcohol avoidance therapy; mechanism involves inhibition of acetaldehyde dehydrogenase, leading to acetaldehyde accumulation and a hangover-like reaction when ethanol is consumed.

  • Mechanism (disulfiram): Ethanol → acetaldehyde (via alcohol dehydrogenase) → acetaldehyde cannot be efficiently converted to acetate due to aldehyde dehydrogenase inhibition by disulfiram, producing adverse effects.

Avoidance therapy – disulfiram example

  • Mechanism: Disulfiram inhibits aldehyde dehydrogenase; ethanol metabolism accumulates acetaldehyde, causing symptoms such as nausea, vomiting, palpitations, dyspnea, flushing.

  • Clinical use: adjunct to rehab to create a negative association with drinking.

Antipsychotics – overview and neurobiology of psychosis

  • Psychosis: loss of contact with reality; symptoms include hallucinations, delusions, emotional blunting, and reduced insight.

  • Schizophrenia: multifactorial etiology (genetic, neurobiological, environmental); structural brain changes and functional alterations are observed.

  • Dopamine hypothesis (core): dysregulation of dopamine pathways contributes to schizophrenia; mesolimbic overactivity linked to positive symptoms; mesocortical hypoactivity linked to negative/cognitive symptoms.

  • Other neurotransmitters (glutamate, serotonin) also implicated through NMDA receptor hypofunction and serotonergic involvement.

Dopaminergic pathways relevant to antipsychotics

  • Mesolimbic pathway: positive symptoms; blockade reduces these symptoms but may reduce reward pathways.

  • Mesocortical pathway: negative and cognitive symptoms; blockade may worsen or fail to improve these symptoms.

  • Nigrostriatal pathway: motor control; blockade produces extrapyramidal symptoms (EPS).

  • Tuberoinfundibular pathway: dopamine regulation of prolactin; blockade can cause hyperprolactinemia (breast enlargement, galactorrhea, reduced libido).

Antipsychotics – major categories

  • Typical (first-generation) antipsychotics: Chlorpromazine, Haloperidol, Droperidol; mainly D2 antagonists with narrower receptor profiles.

  • Atypical (second-generation) antipsychotics: Clozapine, Olanzapine, Risperidone; broader receptor activity including serotonin and histamine receptors; generally lower risk of EPS and hyperprolactinemia, but other side effects exist.

  • Distinction is not absolute, but typicals are more selective for D2 antagonism; atypicals have broader receptor activity.

Actions and adverse effects of antipsychotics

  • Therapeutic action: blockade of D2 receptors in the mesolimbic pathway reduces positive symptoms.

  • Extrapyramidal side effects (EPS): due to D2 blockade in the nigrostriatal pathway; include dystonia, akathisia, parkinsonism, tardive dyskinesia.

  • Endocrine effects: blockade in tuberoinfundibular pathway increases prolactin release; can cause galactorrhea and other sexual dysfunction.

  • Antiemetic effects: dopamine blockade in chemoreceptor trigger zone reduces nausea/vomiting.

  • Antihistamine and antimuscarinic effects: blockade can cause sedation and dry mouth, constipation, blurred vision, tachycardia, urinary retention.

  • Other receptors: blockade of α-adrenergic receptors can cause orthostatic hypotension and other autonomic effects.

Extrapyramidal side effects and time course

  • EPS types:

    • Dystonic reactions (acute muscle spasms), akathisia (restlessness), Parkinsonian symptoms (rigidity, tremor), tardive dyskinesia (late-onset involuntary movements).

  • Risk increases with longer treatment duration and higher D2 occupancy; some antipsychotics carry different EPS risk profiles.

Antipsychotics – practical safety considerations

  • Hyperprolactinemia risk contributes to sexual dysfunction and adherence issues.

  • Sedation and anticholinergic effects can be problematic for elderly patients (falls, cognitive impairment).

  • Atypicals may offer better tolerability for negative symptoms but require monitoring for metabolic effects (weight gain, dyslipidemia, diabetes risk).

Summary connections to foundational principles and real-world relevance

  • CNS structure–function relationships underpin pharmacologic targeting: blocking specific neurotransmitter receptors in particular circuits can relieve symptoms but may introduce adverse effects in other circuits.

  • The balance between dopamine and acetylcholine in motor pathways explains Parkinsonian-like symptoms with some antipsychotics and the rationale for anticholinergic co-therapy in EPS.

  • Multiple neurotransmitter systems (dopamine, serotonin, glutamate, GABA) contribute to complex psychiatric disorders; this underlies the rationale for atypical antipsychotics with broader receptor activity beyond D2 antagonism.

Quick reference: key numerical/temporal points

  • Benzodiazepine use should be limited to approximately 14\text{–}21\text{ days} to minimize tolerance and dependence.

  • Flumazenil is used to reverse benzodiazepine toxicity and has a short half-life, requiring IV administration.

  • Extrapyramidal symptoms are a major concern with typical antipsychotics due to nigrostriatal D2 blockade.

Connections to assessed topics and exam-style concepts

  • Distinguish anxiolytics from hypnotics by dose and CNS effects; understand receptor mechanisms (GABA_A for most benzodiazepines).

  • Recognize the dopamine hypothesis in schizophrenia and how antipsychotics modulate pathways differently, impacting positive vs negative symptoms.

  • Understand the practical pharmacokinetic implications of benzodiazepines (lipophilicity, fat storage, active metabolites) for onset, duration, and withdrawal risk.

Illustrative concepts and examples (hypothetical scenarios)

  • If a patient presents with tremor, rigidity, and slowed movement after starting a typical antipsychotic, consider extrapyramidal side effects linked to nigrostriatal D2 blockade and evaluate dose reduction or switch to an atypical agent.

  • In treating insomnia with a benzodiazepine, anticipate possible daytime sedation or cognitive effects, particularly in older adults; prefer shorter-acting agents when possible and monitor for dependence.

  • Disulfiram as an avoidance strategy creates a strong negative association with alcohol by producing unpleasant symptoms after alcohol ingestion, leveraging learned conditioning to support abstinence.

Links to foundational pharmacology principles

  • GABAergic modulation underpins many sedatives and hypnotics, illustrating how enhancing inhibition can reduce CNS excitability across multiple circuits.

  • Dopaminergic pathways illustrate how regional neurotransmitter activity aligns with symptom clusters (positive vs negative symptoms) in psychiatric disorders.

  • Receptor diversity (D1/D2, 5-HT subtypes, NMDA/mGluR, GABA_A/B) explains variability in drug effects, side effects, and patient responses.

Table-style quick reference (synthesis of Table 32.1 concepts)

  • Brain region associations:

    • Cerebral cortex, thalamocortical tracts, pyramidal pathway, reticular activating system: Acetylcholine; Dopamine involvement in extrapyramidal pathways and the limbic system; Serotonin involvement in arousal and mood.

    • Hypothalamus: Serotonin and Noradrenaline related in hormone release and autonomic control; Dopamine involvement in prolactin regulation via tuberoinfundibular pathway.

    • All regions: GABA and Glutamate as fundamental inhibitory/excitatory transmitters across CNS.

  • Functional roles:

    • Cognition, movement, memory, consciousness: Acetylcholine and Glutamate.

    • Motivation, behavior, vomiting, hormone release: Dopamine, Serotonin, Noradrenaline.

    • Arousal, sleep, mood, appetite, temperature: Serotonin, Noradrenaline, Histamine, Dopamine.