BIOL 1020 Exam 3
Photosynthesis
Process that converts solar energy into chemical energy
Directly or indirectly, photosynthesis nourishes the entire world
Classification by Energy and Carbon Sources
Energy Source
Chemotrophs: can only get energy directly from chemical compounds
Phototrophs: get energy directly from light (these organisms can use chemical compounds as energy sources as well)
4 sub groups
Photoautotrophs: carry out photosynthesis
Photoheterotrophs: use light energy but cannot fix CO2; some non sulfur purple bacteria
Chemoautotrophs: obtain energy from reduced inorganic molecules and use some of it to fix CO2; some bacteria
These were the first cells on earth
Chemoheterotrophs: use organic molecules as both carbon and energy sources
Autotrophs and Heterotrophs
Autotrophs: obtain energy directly from the environment, and making organic molecules from inorganic molecules
Heterotrophs: get energy and carbon by breaking down organic molecules assembled by other organisms
Electromagnetic Spectrum
Molecules can absorb photons, thus becoming energized
Chloroplasts
In photosynthetic eukaryotes, photosynthesis occurs in chloroplasts
Chloroplasts have both an inner and outer membrane
Stroma: fluid-filled region inside the inner membrane
Sugars are built in the stroma (second stage of photosynthesis)
Thylakoids: disklike membranous sacs found in stroma (interconnected with each other and inner membrane)
Thylakoid Membrane: folded inner membrane that makes up thylakoids
Thylakoid Lumen: fluid-filled region inside a thylakoid
Granum: stack of thylakoids
Chlorophyll: main light harvesting molecule, found in the thylakoid membrane
Pigments
Pigment: an organic molecules that selectively absorbs light of specific wavelengths
Reflects wavelengths no absorbed and gives it color
Chlorophyll a
Most common photosynthetic pigment
Absorbs violet-blue and red light
Reflects green
Accessory pigments are also found in the thylakoid membrane
Can transfer captured energy to chlorophyll a
Photosynthesis
H20 is oxidized and CO2 is reduced (endergonic process)
Divided Into:
Light Reactions
Capture light energy
Consume water
Produce oxygen
Energy is placed in ATP and NADPH in the stroma
C3 Cycle
Consumes CO2 and energy (proved by ATP and NADPH)
Produces carbohydrates
In many ways, this is the reverse of aerobic respiration
In the thylakoids light energy is received and water is split providing a source of electrons and protons H+, producing O2 gas as waste and NADP and ATP
C3 Cycle
Carbon fixation (carbon uptake)
Taking carbon atoms from inorganic sources (atmosphere) and using them to produce organic molecules
Builds glucose from CO2
CO2 combines with the 5-carbon compound ribulose 1,5-biphosphate (RuBP)
Carbon reduction
3-PGA is reduced to glyceraldehyde 3-phosphate (G3P) in two steps; in the process, ATP and NADPH are used
From 6 CO2 you get 12 G3P
2 G3P are removed and used to make glucose or fructose (thus 6 carbons leave to make C6H12O6)
Remaining 10 G3P are used to regenerate RuBP
RuBP regeneration
Series of 10 reactions
Rearrange the 10 G3P to form 6 ribulose phosphate molecules
C3 Plants: Photorespiration
Stomata is closed so CO2 cannot enter and O2 cannot leave
Low CO2, high O2
Photorespiration
At high O2 levels, rubisco attaches oxygen to RuBP instead of carbon
One PGA and one glycolate form
Glycolate broken down into CO2 and H2O
C4 Plants
Named because they preface the carbon cycle with an alternate mode of carbon fixation that forms a four-carbon compound as its first product
CAM Plants (Crassulacean Acid Metabolism)
A second photosynthetic adaption to arid conditions has evolved in pineapples, cacti, other plants
Only opens stomata at night to keep from losing water, fixes carbon at night
Forms 4-carbon molecule at night
C4 works by altering the location of initial CO2 fixation
CAM works by altering the time of initial CO2 fixation
Why do cells divide?
Multicellular eukaryotes depend on cell division for…
Development from a fertilized cell
Growth
Repair
Cell division is an integral part of cell life
Cells
Somatic cells (body cells/nonreproductive cells) 2n
Gametes (sex cells/reproductive cells: sperm and egg) n
Prokaryotes reproduce by a type of cell division called binary fission
Usually a singular, circular chromosome
Eukaryotic DNA
Chromatin: long DNA molecule with associated proteins
Chromosomes: densely packaged chromatin
DNSA gives instructions to make protein or RNA molecule
Genome: organism's complete DNA sequence
Mitosis
Human cell might undergo one division in 24 hours
4 Stages
Prophase
Metaphase
Anaphase
Telophase
Prophase
Chromatin condenses to form chromosomes
Each chromosome (duplicated during s phase) forms a pair of sister chromatids
Joined by a centromere
System of microtubules called the mitotic spindle, organizes between the two poles of the cell
By end of prophase, nuclear membrane has broken down and nucleoli have disintegrated
Metaphase
Chromosomes line up along the midplane of the cell (the metaphase plate)
Chromosomes are highly condensed
Mitosis checkpoint appears to be here
Sister chromatids separate and are moved toward opposite poles
Anaphase
Sister chromatids separate and are moved toward opposite poles
Motor proteins move the chromosomes towards the poles along the kinetochores microtubules
Overall, this process assures that each daughter cell will receive one of the duplicate sets of genetic material carried by the chromosomes
Telophase
Prophase is essentially reversed
Mitotic spindle is disintegrated
Chromosomes decondense
Nuclear membranes reform
Cytokinesis
Divides the cell into two daughter cells
Usually beings in telophase and ends
In animals, a cleavage furrow develops
In plants, a cell plate develops
Cytoplasm is usually distributed randomly but roughly equally between daughter cells
Cell Signaling
Endogenous: cells can receive signals to divide from within the cell
Exogeneous: cells can also be signaled to divide by chemical factors outside the cell
Asexual Reproduction
Creates offspring that are genetically identical to each other and to the parent cell (clones)
Only mitotic cell division is required
Typically rapid and efficient compared to sexual reproduction
Sexual Reproduction
Occurs when specialized sex cells called gametes fuse to form a single cell called a zygote
Offspring are not genetically identical to their parents
Genetic recombination
May produce some offspring that are better adapted to the environment than either parent
May produce some offspring that are moor poorly adapted than either parent
Must have a way to half the number of chromosomes at some point, otherwise the number of chromosomes would double with each generation
Accomplishes through meiosis
Diploid cells give rise to haploid cells during meiosis
Most somatic cells are diploid
Diploid (2n): two complete sets
Haploid (n): one set
Polyploid (3n or more): extra sets
Three events are unique to meiosis and all occur in meiosis I
Synapsis and crossing over in prophase I
At metaphase I --> paired homologous chromosomes (tetrads)
At anaphase I, homologous chromosomes, NOT sister chromatids separate
Genetic variation in sexual life cycles contributes to evolution
Mutations are the original source of genetic diversity
Mutations create different versions of genes called alleles
Reshuffling alleles during sexual reproduction produces genetic variation
Crossing Over
Produces recombinant chromosomes, which combine DNA inherited from each parent
Crossing over begins very early in prophase I, as homologous chromosomes pair up gene by gene
In crossing over, homologous portions of two non-sister chromatids trade places
Crossing over contributes to genetic variation by combining DNA from two parents into a single chromosome
Independent Assortment
Homologous pairs of chromosomes orient randomly at metaphase I of meiosis
Random Fertilization
Any sperm can fuse with any ovum