Lecture 15: Competition 1 - October 13th

Course Announcements and Housekeeping

  • Good Morning: Instructor checks in with students after a break.

  • Exam 2: Scheduled for next Wednesday (not this Wednesday).

    • Resubmissions for Exam 1: Will be checked today until Wednesday.

    • Exam Key: Will be posted after resubmissions, allowing students to check their answers with the final grading.

    • Corrections Timeframe: Students have two weeks after the exam to discuss any grading issues.

  • Final Paper Discussion:

    • Specifics about the final paper will be addressed, but the discussion will not dominate today's lecture, which focuses on the upcoming exam and the topic of interspecific interactions.

    • No final in-person exam; students will simply submit this final paper for their grade.

Upcoming Lecture Topics

  • Main Focus: Interspecific interactions, particularly competition.

  • Lecture Schedule:

    • Today: Competition and possibly predation (if time permits).

    • Following lectures will continue exploring these topics leading up to Exam 2.

Competition in Ecological Context

  • Interspecific vs. Intraspecific Competition: Highlighted importance of differentiating between these two types.

    • Intraspecific Competition: Discussed in past lectures; refers to competition among individuals of the same species.

    • Interspecific Competition: Focus of today's lecture, referring to competition among different species.

Understanding Niche Concepts

  • Niche Definitions:

    • Fundamental Niche: Theoretical space with many dimensions representing environmental conditions that allow survival and reproduction for a species.

    • Realized Niche: Actual space occupied by a species, reduced by competition with other species.

    • Comparison:

    • The fundamental niche includes all potential resources a species could utilize in an ideal situation.

    • The realized niche accounts for limitations imposed by the presence of competing species.

Learning Goals

  • Students should:

    • Understand definitions of both types of niches.

    • Learn how to measure and relate these concepts to others previously covered.

    • Recognize links to demographic processes such as survival, reproduction, and population growth.

Competition Dynamics

  • Physiological and Biotic Filters: Affect the realization of the niche.

  • Effects of Resources: Resources can facilitate population growth, contrasting with filters that limit populations.

  • Key Graphs & Illustrations: Concepts will be emphasized through graphs and models to differentiate between fundamental and realized niches visually.

Detailed Niche Definitions

  • Fundamental Niche Definition: An imaginary space with many dimensions where each axis represents a range of environmental conditions that allow species survival, growth, and reproduction.

  • Realized Niche Definition: Space representing resources available reduced due to competition with other species.

Intraspecific Competition Dynamics

  • Population Growth: Changes due to competition among individuals of the same species.

  • Logistic Growth Model: Describes how populations grow and stabilize over time, incorporating carrying capacities (k).

    • An observed growth pattern showcasing initial exponential growth that eventually levels off due to resource limitations and competition.

Fundamental vs. Realized Niche Relationship

  • Impact of Intraspecific Competition on Realized Niche: Intraspecific competition does not change how a species exploits resources but affects population regulation.

  • Exam Question: Does intraspecific competition affect the realized niche?|NO, it doesn’t affect the imaginary space of resource exploitation for a species.|\

Understanding Trade-offs in Ecological Niche

  • Key Concept: A more nuanced understanding reveals that species do adapt their niches based on competition, leading to morphologically, physiologically, and behaviorally inspired adaptations over evolutionary time.

Interspecific Competition Evaluation

  • Two Species Interaction: When two species compete, they exert negative effects on one another's growth and reproductive success.

  • Measurement of Effects: Reduction in fecundity, survivorship, and/or growth noticed when two different species interact.

Laboratory vs. Natural Examples of Competition

  • Gauss (1934) Paramecium Experiment: Laboratory demonstration showing that when competing species of protists were placed together, one would eventually dominate and lead to exclusion of the other.

    • Graphs Analyzed: Display separate growth rates of species when grown alone versus together, illustrating interspecific competition effects.

Principle of Competitive Exclusion

  • Definition: States that when two species are placed together competing for the same resource, one will always outcompete the other.

  • Coexistence in Varying Contexts:

    • Natural Species Instantiation: Example with mongoose species showcasing ecological release when one competitive pressure removed leads to increased size parameters in the less dominant species.

Real-world Example of Niche Adaptations

  • Natural Selection in Action: Through the presence of competitors, species will shape their morphological traits accordingly to minimize competition.

  • Character Displacement vs. Ecological Release: Focusing on species adaptation dynamics, where removal of competition leads to optimal ecological niche realization over time.

Concluding Remarks and Next Steps

  • Summary of Interspecific vs. Intraspecific Competition: Types of competition with illustrations.

  • Next Class Focus: Move deeper into understanding interspecific competition dynamics and applicable examples, including further lab experiments to define coexistence and resource partitioning among competitors.