Evolution Notes

Evolution

  • Change over time.
  • Visual representation of Apple products from 1976-2007 showing form factor evolution.

Geologic Time Scale

  • Covers 650 million years to the present.
  • Illustrates the change in DNA (mutations) over time, directed by natural selection.
  • Changes occur over many generations.
  • Key Eras, Periods, and Events:
    • Cenozoic Era:
      • Quaternary Period (0-2.6 million years ago): Evolution of humans.
      • Neogene Period (2.6-5 million years ago): Mammals diversify.
      • Paleogene Period (5-50 million years ago): Extinction of dinosaurs, first primates.
    • Mesozoic Era:
      • Cretaceous Period (50-150 million years ago): First flowering plants, first birds.
      • Jurassic Period (150-200 million years ago): Dinosaurs diversify.
      • Triassic Period (200-250 million years ago): First mammals, first dinosaurs.
    • Paleozoic Era:
      • Permian Period (250-300 million years ago): Major extinctions, reptiles diversify.
      • Carboniferous Period (300-350 million years ago): First reptiles, scale trees, seed ferns.
        • Pennsylvanian.
        • Mississippian.
      • Devonian Period (350-400 million years ago): First amphibians, jawed fishes diversify, first vascular land plants.
      • Silurian Period (400-450 million years ago).
      • Ordovician Period (450-500 million years ago): First fishes.
      • Cambrian Period (500-550 million years ago): Sudden diversification of metazoan families, first chordates, first skeletal elements, first soft-bodied metazoans, first animal traces.
    • Late Proterozoic Era: (550-650 million years ago).

Lamarck vs. Darwin's Theories of Evolution

  • Lamarckism contrasted with Darwinism.

Lamarck’s Theory

  • Inheritance of acquired characteristics.
    • Individuals acquire traits during their lifetimes due to experience or behavior.
    • These traits are then passed to offspring.
    • Example: Giraffes evolved long necks by stretching to reach leaves, passing longer necks to offspring.
  • Proven wrong by Weismann in the 1870s.
    • Acquired traits are not genetically determined.

Darwin’s Theories

Charles Darwin

  • British naturalist (1809-1882).
  • Observed differences in species across geographic regions, particularly in the Galapagos Islands.

Darwin’s First Theory: Descent with Modification

  • Fossils are similar to, but different from, living organisms.
  • Descent with Modification:
    • Newer forms in the fossil record are modified descendants of older species.
    • This is a result of random genetic mutations.

Darwin’s Second Theory: Natural Selection

  • Mechanism for change in a population.
  • Causes evolution in nature over millions of years.
  • The environment selects favorable traits for success and reproduction.

Environment as a Driving Force

  • Environments on Earth are constantly changing:
    • Gradual or sudden changes in climate, human activity, natural disasters, etc.
    • Migration can also change the environment for an organism.

Genetic Variations

  • Organisms in a population have differences.
  • Biodiversity is crucial.
    • Higher genetic diversity reduces the likelihood of extinction in a changing environment.

Sources of Variation

  • Mutations: flawed copies of genes (related to Darwin’s Descent with Modification).
  • Recombination: independent assortment of genes and crossing over in meiosis.
  • Sexual Reproduction: Random fusion of gametes between two individuals.
  • Environment: plays a role; less diversity increases extinction risk.

Overproduction of Offspring

  • More organisms are produced than will survive.
  • This allows for a greater chance of survival for the species.

Struggle for Survival/Competition

  • Individuals compete for food, mates, and space.
  • Individuals with favorable variations are more successful, survive, and reproduce (survival of the fittest).

Survival and Reproduction

  • Favorable variations (adaptations) are passed on.
  • Unfavorable variations disappear.
  • Individuals that survive are the most fit or best suited for their environment (adapted).

Natural Selection Leads to Adaptations and Fitness

  • Adaptation: Result of natural selection.
    • Inherited variation (favorable trait) that increases a population’s chance of survival in a given environment.
    • Examples: physical traits, behaviors, biochemical processes.
    • Galapagos finches: beak adaptations for specific environments.
  • Fitness: An organism naturally selected by the environment is adapted or better suited for survival.

Questions on Natural Selection

  • What drives natural selection?
    • The environment selecting favorable traits.
  • Is natural selection an active process?
    • No, traits are inherited, not acquired.
  • Alternative to natural selection?
    • Artificial selection/selective breeding (not natural).

Artificial Selection/Selective Breeding

  • Darwin’s theory is based on evidence of artificial selection.
  • Producing variations of the same species by selecting for desired traits in offspring.
  • Examples: pigeons, dogs, crops, cattle.

Population Genetics - Evolution at the Genetic Level

  • A group of individuals of the same species living in the same area and sharing a common gene pool.
  • Gene pool: The sum of all genetic information (genes) carried by members of a population.

Population Genetics

  • Individuals do not evolve; populations do.
  • Evolution is the gradual change of allele frequencies in a population (microevolution).
  • Gene/Allele frequency: Percentage of an allele in a gene pool.
  • Favorable genes become more frequent.
  • Example: Rock pocket mouse frequency changed from d to D with environmental change.

Hardy-Weinberg Genetic Equilibrium

  • Allele frequencies in a population remain the same over generations unless acted on by outside influences.
  • Populations do not evolve but remain constant under specific conditions.

Equilibrium Conditions

  • No mutations
  • No migration
  • Large population size
  • Random mating
  • No selection (natural or artificial)

Real Populations vs. Theoretical

  • Real populations violate conditions necessary for genetic equilibrium.
  • This leads to variations among organisms, natural selection, and evolution.

5 Processes of Evolution

  • Allele frequencies remain constant unless outside forces are at play.
  • These processes cause allele frequencies/evolution to occur.

1. Genetic Drift

  • Random change in allele frequency due to chance.
  • Small populations (<100) most affected.
  • Alleles tend to become lost or fixed.
  • Examples: Natural disasters, new arrivals to an isolated area.

2. Nonrandom Mating

  • Mate selection influenced by geographic proximity or similar characteristics.

3. Mutation

  • Random inheritable change in genetic material.
  • Can cause new alleles or forms of a trait.
  • Can be harmful, fatal, neutral, or beneficial in a changing environment.

4. Migration

  • Movement of individuals in or out of a population.
  • Gene flow: sharing of genes through sexual reproduction.
    • Consistent gene flow maintains species.
    • Immigration enhances variation (speciation), emigration decreases it.
  • Also referred to as geographic isolation.

5. Natural Selection

  • Most fit survive and pass down traits.
  • Acts on existing genetic variation.
  • Favorable traits are selected by environment.
  • Results in adapted organisms (fit).
  • Can lead to extinction of others (macroevolution).

Types of Selection

Stabilizing Selection

  • Favors average phenotypes over extremes.
  • Example: Medium height plants are favored because short plants can't compete for sunlight and tall plants are susceptible to wind damage.

Directional Selection

  • Favors phenotypes at one extreme.
  • Example: Giraffe necks - selection pressure against short necks resulted in a shift towards longer necks.

Disruptive Selection

  • Takes place when extremes of phenotypic range are selected.
  • Eliminates the average.
  • Rare.
  • Example: Plant pollinated by three different pollinators that prefer short, medium and tall plants. If the pollinator for medium plants disappears the population favors short and tall plants, resulting in a polymorphic population.

Evolution in Modern Times

  • Antibiotic resistance is an example affecting lives today.
    • What is this?
    • What has contributed to this problem?

Examples of Evolution in Action

  • Bacterial resistance to antibiotics
  • Mosquito resistance to insecticides
  • Peppered moth changes with the industrial revolution