Evolution and Characteristics of Animals
Defining Animals
Characteristics of Animals:
Heterotrophs: Organisms that obtain their food by consuming other organisms.
Multicellular: Composed of multiple cells.
No cell walls: Unlike plants, animals do not have rigid cell walls.
Movement: All animals have some form of movement at some point in their life cycle.
Diversity in morphology: Variation in shape and structure among different species, influenced by HOX genes.
Diversity in habitats: Animals can live in various environments, including terrestrial, aquatic, and aerial.
Sexual Reproduction: Most animals reproduce sexually, involving the combination of genetic material from two parents.
Tissues: Animals are composed of specialized cells forming tissues.
Evolution of the Animal Body Plan
Five Key Innovations in the evolution of animal body plans:
Tissues
Symmetry
Body cavity
Development
Segmentation
Evolution of the Animal Body Plan
Evolution of Tissues:
Parazoa: Organisms that lack defined tissues and organs. Example: Phylum Porifera (sponges).
Eumetazoa: Organisms with distinct tissues and irreversible differentiation for most cell types, encompassing all other animals.
Evolution of Symmetry:
Parazoa: Lack symmetry.
Eumetazoa: Display symmetry along an axis through the body.
Types of Symmetry:
Radial Symmetry: Organisms are arranged around a central axis, allowing two equal halves in any 2-D plane.
Bilateral Symmetry: Body can be divided into right and left halves that are mirror images. The sagittal plane bisects the body into two equal halves.
Significance: Symmetry enhances complex movement capabilities.
Advantages of Bilateral Symmetry:
1. Cephalization: Development of a cranial central nervous system (CNS).
2. Greater Mobility: Beyond linear movement.
Evolution of Coelom (Body Cavity):
Eumetazoa: Possess three germ layers:
Ectoderm: Forms body coverings and nervous system.
Mesoderm: Develops into the skeleton and muscles.
Endoderm: Forms digestive organs and intestines.
Coelom: A space surrounded by mesodermal tissue formed during development.
Types of Body Cavities:
Acoelomates: Organisms lacking a body cavity.
Pseudocoelomates: Body cavity located between mesoderm and endoderm.
Coelomates: Body cavity completely enclosed in mesoderm.
Patterns of Development in Bilaterians: Two groups:
Protostomes: Develop mouth first from the blastopore, anus (if present) develops either from the blastopore or another region of the embryo.
Deuterostomes: Develop anus first from the blastopore, mouth later from another region of the embryo.
Segmentation:
Definition: The division of the body into regions, allowing for more efficient and flexible movement.
Functions: Development of body regions for different functions and regeneration of body parts in some taxa.
Evolutionary Appearance: Segmentation has appeared several times in animal evolution.
The Earliest Animals
Dickinsonia: An organism from the Ediacaran biota (570 - 550 million years ago), exhibited bilateral symmetry and suggested evidence of cholesterol.
Charniodiscus: Another significant organism from the same era.
Animal History
Cambrian Period: By 500 million years ago, all major animal phyla were present.
Cambrian Explosion: A major adaptive radiation event that occurred over 30 million years, significantly evolving basic shapes and sizes.
Burgess Shale: Key fossil site for studying this evolutionary change.
Homeobox (Hox) gene complex: This genetic framework evolved during this period, leading to rapid changes in animal body plans.