Population Ecology

POPULATION ECOLOGY

Levels of Organization

  • Global ecology: Focus on global interactions between ecosystems.
  • Landscape ecology: Examines spatial patterns and how they affect ecosystems.
  • Ecosystem ecology: Studies energy flow and nutrient cycling within ecosystems.
  • Community ecology: Analyzes interactions between different species in a community.
  • Population ecology: Investigates factors affecting population size and distribution of a species.
  • Organismal ecology: Looks into individual organisms and their adaptations to the environment.

Ecology: All Life is Interconnected

  • What do I need to know?
    • Major structural and dynamic characteristics of populations.
  • What do I need to think about?
    • Populations have the potential for exponential growth but rarely realize it.
  • What do I need to understand?
    • How life history traits provide tools to "Win the Game" or maximize growth in species like zebra mussels.
    • Understanding of dependent and independent factors affecting populations.

Population Ecology

  • Definition: Population ecology explores the biotic and abiotic factors that influence density, distribution, and size of populations.
  • Key Concepts:
    • Population: A group of individuals of a single species living in the same general area.
    • Described by boundaries and size.
    • Characterized by density, dispersal, and demographics.

Density

  • Definition: The number of individuals per unit area or volume.
  • Dynamic Factors:
    • Births: Introduce new individuals into a population.
    • Deaths: Remove individuals from a population.
    • Immigration: Individuals moving into a population.
    • Emigration: Individuals moving out of a population.

Patterns of Dispersion

  • Dispersion: A pattern of spacing among individuals within the boundaries of the population.
    • Types of Dispersion:
    • (a) Clumped: Individuals are grouped in patches.
    • (b) Uniform: Individuals are evenly distributed.
    • (c) Random: Individuals are spaced in an unpredictable manner.

Demography

  • Definition: Study of the vital statistics of a population and how they change over time.
  • Vital Statistics:
    • Survivorship: The proportion of individuals surviving at different ages.
    • Reproductive rate: Includes:
    • Fecundity: The number of offspring potentially produced per reproductive cycle.
    • Fertility: The actual number of offspring produced.

Survivorship Curve

  • Definition: A graphical representation of data in a life table.
  • Types of Survivorship Curves:
    • Type 1: High survivorship until old age.
    • Type 2: Constant mortality rate regardless of age.
    • Type 3: High mortality in early life but survivorship increases for those who reach a certain age.

Reproductive Rates

  • Reproductive table: An age-specific summary of reproductive rates of a population.
    • Typically a snapshot of the population at a specific time, focusing on the total female reproductive output (fertility).

Population Growth

  • Function of four variables:
    • Births, deaths, immigration, emigration.
  • Closed Population: A population with no immigration or emigration.
  • Growth Rate:
    • Change in population size can be pronounced as:
      \text{Change in population size} = \text{Births} + \text{Immigrants} - \text{Deaths} - \text{Emigrants}

Population Growth Equation

  • Basic Equation for Growth Rate (r): \Delta N / \Delta t = rN
    • Where r = b - d (b = birth rate, d = death rate).

Exponential Growth

  • Definition: Growth under idealized conditions where the rate of increase is maximum (rmax).
  • Equations:
    • \frac{dN}{dt} = r_{max}N
    • Example: 1.0N, or 0.5N depending on conditions.

Example: Elephant Population

  • Location: Kruger National Park, South Africa post-hunting ban observed exponential growth in elephant population leading to:
    • Elephant population increase from years 1900 – 1970 observed in the graphical representation.

Example: Zebra Mussels

  • Characteristics of Zebra Mussels:
    • Highly invasive species with a max size of 50mm.
    • Age of first reproduction: 2 years; Lifespan: 4-5 years.
    • Fecundity/spawn: 40,000 eggs.
    • Filtration capacity: 1 liter algae/day.
  • Invasion Routes: Hitchhiked from Europe via ship ballast to Great Lakes, particularly Lake St. Clair in the U.S.

Ecological Impacts of Zebra Mussels

  • Impact on Native Species:
    • Loss of native mussels, alteration in energy flow, enhanced water clarity.
    • Increased growth in aquatic plants, effects on fish communities (not fully studied).

Carrying Capacity

  • Definition: The maximum population size that an environment can support.
  • Concept: Exponential growth cannot be sustained indefinitely due to limiting factors.

Logistic Growth Model

  • Definition: Describes growth rate declines as carrying capacity is approached.
  • Equation: \frac{dN}{dt} = rN\left(\frac{K - N}{K}\right)
    • Produces a sigmoid (S-shaped) growth curve.

Logistic Growth Example

  • Hypothetical Population: $K = 1500$. Growth rate per capita varies as the population size approaches carrying capacity.
  • Calculation: Demonstrated through a table showing population size vs. growth rate.

Real Populations vs Logistic Growth

  • Phenomenon Observations:
    • Natural populations often overshoot carrying capacity and stabilize through high mortality (often termed as ‘compensation’).

Life History Traits

  • Definition: Traits affecting reproduction and survival schedules of organisms.
  • Key Traits Include:
    • Age of maturity, reproductive frequency, and number of offspring.

Trade-offs in Life History Strategies

  • Maximization Strategy Characteristics:
    • Fast growth rate, early maturation, high fecundity, larger offspring size.
  • Types of Strategists:
    • R-Strategists: Live fast, reproduce often, no care for offspring.
    • K-Strategists: Live slowly, care for fewer offspring, mature later.

Population Regulation Mechanisms

  • Density-independent regulation: Not affected by population size (e.g., abiotic factors).
  • Density-dependent regulation: Affected by biotic factors such as competition, predation, and disease.

Dynamics of Population Interactions

  • Focus: On complex interactions. Nature is influenced by multi-variable conditions, incorporating both density-dependent and independent factors.