Signal Transduction and Second Messengers
Module 4: Signal Transduction and Second Messengers
By the end of this module, you should be able to:
Describe the different ways cells can communicate with each other.
Gap Junctions: connexin proteins make little ion-carrying tunnels between cells called connexons, and control the rapid and direct ion passage
Cell-to-cell binding: surface molecules bind together
Extracellular messengers:
Hormones: Extracellular messengers carried by blood
Neurotransmitters: released from a neuron into the blood where it then acts similar to hormones
Local mediators: Released by cells and act locally on other cells
Autocrine = shuts off signal
Paracrine = turns on signal
3 types:
Cytokine = cell growth
Nitric oxide = causes relaxation of blood vessel (dilation)
Eicosanoids = released after stimuli
Prostaglines
Leukotrienes
Thromboxane
Compare and contrast the signal transduction mechanisms for lipophilic and lipophobic messengers.
Describe how extracellular messengers reach their target cells.
Depends on type:
Hormones: through the blood
Neurotransmitters: through synapse
Discuss the properties of messenger-receptor binding.
Receptor: protein on the cell, accepts or denies
There can be one receptor for many messengers
There can be many receptors that accept the same messenger
Which receptor uptakes it depends on the affinity
*ones with higher affinity to the messenger will take priority
Explain how receptors are regulated.
Upregulations: messenger deficiency = the cells may increase the number of receptors.
Downregulation: excess messengers = cell receptor number decreases
Identify the components of a signal transduction pathway and explain how the pathways are amplified.
The sequence of events between receptor activation & the cell’s response
Extracellular (first) messenger binds to receptor
Open or close channel
Activate intracellular (second) messenger system
G protein activated
Adenyl cyclase activated
Generates hundreds of cAMP molecules
Activates protein kinase A
Phosphorylated hundreds of proteins
(each step multiplies the responders)
Terminated by protein degradation, diffusion of messenger, removal of calcium from cytosol and engulfing the complex
Describe the following functional classes of receptors:
Channel-linked receptors: Simplest receptors. Messenger binds → channel opens (ex: Ca++ channels)
Enzyme-linked receptors: Messenger binds → receptor activates attached enzyme → catalyzes chemical reaction (tyrosine kinase… think growth factors)
G protein-linked receptors: Largest category. G protein binds to a GDP or GTP. “switch”(receptor communicating to ion channels or enzymes).
Independent of a second messenger.
Direct communication between G proteins to open OR close ion channels. (slow gating)
Indirect communication - G protein cAMP → phosphorylate plasma membrane channel = open/close