Water Beetles and Backswimmers
Water beetles can extract oxygen from water, maintaining a positive buoyancy.
Backswimmers utilize hemoglobin to maintain neutral buoyancy by minimizing gas exchange with water.
They initially rely on gas in their trachea for buoyancy, utilizing oxygen when needed to achieve neutral buoyancy, which conserves metabolic energy.
Fick's Law of Diffusion
J (diffusion rate) can be increased by increasing surface area, essential for larger organisms.
Air contains ample oxygen (21%) allowing for thicker membranes for gas exchange compared to aquatic environments.
Surface Area in Human Lungs
Each lung contains approximately 300 million alveoli, creating a surface area around 75 square meters (roughly the size of a tennis court).
Thoracic Cavity Dynamics
Inhalation involves contraction of external intercostal muscles, increasing thoracic cavity volume and decreasing pressure, causing air to rush in.
Exhalation involves relaxation of the external intercostals and contraction of internal intercostals to compress the thoracic cavity and increase pressure, expelling air.
Air Flow Dynamics
Humans exhibit tidal flow of air, where air enters and exits through the same opening, leading to mixing of fresh and stale air, creating dead air space.
The inclusion of dead air makes breathing less efficient, as not all inhaled air is fresh.
Lung Volume Definitions
Inspiratory Reserve Volume: Air volume beyond normal inhalation.
Expiratory Reserve Volume: Air volume beyond normal exhalation.
Vital Capacity: The total amount of air moved in and out, excluding residual air that remains in the lungs.
Residual Volume: Air that never gets expelled, contributing to dead air space.
Gas Exchange Efficiency
During tidal breathing, oxygen partial pressure drops from 100 mmHg to 60 mmHg when traveling to the capillaries due to dead air space, reducing efficiency.
The actual difference between inhaled air and the blood reaching the lungs represents the usability of oxygen in the body.
Comparison of Breathing Mechanisms
Birds and Fish: Exhibit unidirectional flow of air or water, enhancing gas exchange efficiency compared to tidal flow.
Concurrent Exchange: Blood and media flow in the same direction; maximal exchange is only 50% due to the gradient diminishing quickly.
Countercurrent Exchange: Blood flows against the media, maintaining a continuous gradient for gas diffusion.
Cross-current Exchange: Blood encounters medium with oxygen at varying concentrations across the branches, achieving better efficiency than concurrent but lesser than countercurrent.
Exchanging Mechanisms
Concurrent Flow: Limited to 50% exchange efficiency.
Countercurrent Flow: Maximizes exchange efficiency sustaining the gradient.
Cross-current Flow: Intermediate efficiency, as blood always initially contacts low oxygen media before moving along.