HO - 03.2 Membranes
Cells and Cell Membranes
Living organisms are composed of one or more cells.
Categories of cells:
Unicellular: Single-celled organisms (e.g., single-celled protists).
Multicellular: Organisms composed of multiple cells.
Overview of Cell Membranes
Plasma Membrane: Defining boundary of the cell.
Structure and types: animal cell plasma membrane and human RBC plasma membrane.
Number of Cells
Unicellular Organisms: Example of a single-celled protist.
Colonial Forms: Few cells forming a colony (e.g., Anabaena sp., a filamentous photosynthetic bacteria).
Multicellular Organisms: Composed of many cells.
Example: Human body contains approximately 70 trillion cells.
Cell Size and Multicellularity
Large organisms are made of many small cells instead of a few large cells.
Limitations of cell size: as cells grow, surface area to volume ratio decreases, impacting nutrient transport and waste removal.
Surface Area to Volume Ratio
Surface-area-to-volume problem as a reason for multicellularity.
Calculating Surface Area and Volume:
For a cubic structure:
Surface area increases with dimensions.
Volume increases cubically.
Effects on Transport: Importance of adequate surface area for nutrient and waste transport.
Examples of cell specialization:
Plants have multiple cell types (about 30 types).
Humans contain close to 200 different types of cells.
Properties of the Plasma Membrane
Basic requirements for a cell membrane:
Must contain the cell's contents.
Must be flexible.
Must allow transport in and out of the cell.
Plasma Membrane Structure:
Composed of phospholipid bilayers.
Membrane thickness: approximately 100 nm.
Spontaneity of vesicle formation in water.
Phospholipid Bilayer
Characteristics:
Flexible and not held together by covalent bonds.
Fluid nature allowing lateral movement and rotation of components.
Rare flip-flop motion of lipids.
Membrane Fluidity
Factors affecting membrane fluidity:
Presence of unsaturated fatty acids enhances fluidity.
Saturated fatty acids make membranes less fluid.
Cholesterol in animals regulates fluidity: stiffens at higher temperatures, prevents solidifying at lower temperatures.
Membrane Proteins
Types of membrane proteins:
Integral Proteins: Span the phospholipid bilayer.
Peripheral Proteins: Attached to integral proteins.
Functions:
Transporters: Assist molecules through the membrane (active or passive transport).
Anchors: Facilitate attachment of other molecules to the membrane.
Additional functions include enzymatic activity and signaling.
Carbohydrates in Membranes
Glycolipids and glycoproteins play protective roles.
Surface molecules can have antigenic properties.
Antigen-Antibody Interactions
Antibodies target specific antigens on cell surfaces.
Example with RBCs: contain protein hemoglobin (transports O2 and CO2).
RBC surface sugars identify cells and can include antigens (A, B, H).
Blood Types and Antigens
Blood group classification based on membrane sugars.
Alleles:
O: recessive, encodes non-functional enzyme.
A/B: co-dominant, encode specific transferase enzymes.
Glycolipids serve as antigens for blood groups (e.g., H, A, B).
Prokaryotic Cell Membranes
Structure of prokaryotic plasma membranes:
Comprised of a phospholipid bilayer with transport proteins.
Peptidoglycan layer and outer membrane analysis in future lectures.
More membrane correlates with increased import, export, respiration, photosynthesis, etc.
Internal Membranes and Infoldings
Infolds of plasma membranes (e.g., thylakoids in cyanobacteria) may be:
Continuous with the plasma membrane (e.g., aerobic bacteria respiratory membranes).
Importance of surface area in eukaryotic internal membranes (rough/smooth ER, Golgi apparatus).
Increased Surface Area for Absorption
Outfolds of the plasma membrane enhance absorption capacity (e.g., intestinal epithelial cells).
Review Questions (Examinable Content)
Define unicellular and multicellular organisms.
Estimate the number of cells and cell types in the human body.
Discuss the size limitations of cells.
Identify major molecules constituting the plasma membrane and their functions.
Explain the basis of blood types and the differences between animal and plant cell membranes.
Describe the mechanisms enabling cells to have extra membranes.