Evolution Notes

Charles Darwin

  • English naturalist and author of "On the Origin of Species" (1809-1882).

  • Introduced evolution by natural selection.

  • Naturalism: belief that all organisms and events arose from natural processes.

  • Darwinian Evolution: Descent with modification by natural selection.

Modern Evolution

  • Theory of Evolution includes population genetics, natural selection, and genetic variation (mutations).

  • Natural Selection: Organisms with traits suited to their environment pass on favorable traits.

Classification and Taxonomy

  • Biologists classify organisms using a logical system.

  • Taxonomy: Classifies organisms and assigns universally accepted names.

  • Carolus Linnaeus (1707-1778): Classification based on structural similarity.

    • Placed organisms into groups and assigned scientific names.

  • Binomial nomenclature: Two-part scientific name (genus and species).

Binomial Nomenclature Rules

  • Scientific name: genus (capitalized) and species (not capitalized).

  • All scientific names are in Latin, italicized or underlined.

  • Organisms in the same species can mate and produce fertile offspring.

Phylogeny and Phylogenetic Trees

  • Modern taxonomy considers an organism's phylogeny (evolutionary history).

  • Phylogenetic trees show evolutionary relationships.

  • Branch points indicate the most recent common ancestor.

Large-Scale Processes Influencing Life

  • Continental Drift: Continents move over time, impacting habitats and climates.

  • Mass Extinctions: Large numbers of species become extinct.

  • Adaptive Radiations: Organisms form new species to fill vacant ecological niches.

Population Genetics

  • Population: Group of same species in same area.

  • Population size changes based on births, deaths, immigration, and emigration.

  • Growth Rate: Amount of population size change over time.

Population Growth Dynamics

  • Exponential Growth: Rapid growth with abundant resources and no predators.

  • Logistic Growth: Growth slows as resources become limited, approaching carrying capacity.

  • Carrying Capacity: Number of individuals an environment can support.

Genetic Variance and Speciation

  • Genetic variance: Diversity of genes and traits within a population.

  • Higher genetic variance increases ability to adapt.

  • Speciation: Splitting of one lineage into two or more distinct groups.

Genetic Drift and Gene Flow

  • Genetic Drift: Random change in gene frequency within a population.

  • Gene Flow: Change in gene pool through migration.

Origin of Life and Biogenesis

  • Biogenesis: All living things come from pre-existing living things.

  • Francesco Redi: Disproved spontaneous generation (abiogenesis) with experiments on maggots.

  • Louis Pasteur: Further disproved spontaneous generation.

Abiotic Synthesis of Organic Molecules

  • Scientists hypothesize simple cells formed via abiotic synthesis.

  • Stages include: abiotic synthesis of monomers, formation of macromolecules, packaging into protobionts, origin of self-replicating molecules.

Oparin Hypothesis

  • Proposed early Earth atmosphere contained ammonia, hydrogen gas, water vapor, and hydrocarbons.

  • These gases formed organic compounds like amino acids in water bodies.

Miller-Urey Experiment

  • Tested Oparin's hypothesis, producing organic compounds (amino acids) from gases and electric sparks.

  • Demonstrated abiotic synthesis of organic molecules is possible.

Formation of Cell-Like Structures

  • Microspheres: Spherical; protein molecules organized as a membrane.

  • Coacervates: Droplets composed of different molecules including amino acids and sugars.

First Life-Forms and Protobionts

  • Protobionts: Collections of molecules surrounded by a membrane-like structure.

  • Exhibit reproduction and metabolism; precursors to prokaryotic cells.

RNA as First Genetic Material

  • First genetic material likely RNA due to its variety of shapes and catalytic abilities (ribozymes).

  • Ribozymes: RNA molecules that act as enzymes and can replicate themselves.

Early Cellular Development

  • RNA molecules replicated and responded to natural selection within protobionts.

First Single-Celled Organisms

  • Early cells were anaerobic prokaryotes.

  • Autotrophs evolved, including chemosynthetic organisms.

Rise of Photosynthesis

  • Photosynthetic cells used carbon dioxide and sunlight to produce organic compounds.

  • Damaging byproduct was oxygen, which led to cellular respiration.

Formation of Ozone Layer

  • Oxygen in atmosphere formed ozone layer, protecting Earth from UV radiation.

Rise of Eukaryotes

  • Eukaryotic cells emerged with DNA organized into chromosomes and membrane-bound organelles.

Endosymbiosis Theory

  • Small aerobic prokaryote lived inside larger anaerobic prokaryote.

  • Aerobic prokaryote became mitochondria.

  • Photosynthetic cyanobacteria became chloroplasts.

Evidences Supporting Endosymbiotic Theory

  • Chloroplasts and mitochondria:

    • replicate independently.

    • contain own DNA (circular).

    • have own ribosomes (similar to prokaryotes).

    • have inner membranes containing enzymes found in prokaryotic plasma membranes.

Multicellular Eukaryotes

  • Single-celled eukaryotes gave rise to multicellular forms.

  • Fossil evidence dates back 565 million years.

Morphology and Homologous Structures

  • Morphology: Classification based on structures possessed by the organism.

  • Homologous structures: Same structure, different functions derived from common ancestor.

Analogous and Vestigial Structures

  • Analogous structures: Similar in function but not in structure; not from common ancestor.

  • Vestigial structures: Reduced in size or function from previous ancestor.

Cladistics

  • Method using shared derived characters to establish evolutionary relationships.

  • Derived character: Evolved only within groups under consideration.

Cladograms

  • Diagram showing evolutionary relationships among organisms.

Cellular and Biochemical Similarities

  • Similar cell structures serve as evidence of common ancestry.

  • Similar chemical compounds found within cells show relationships.

Genetic and Embryological Similarities

  • Similarities in number and type of chromosomes.

  • Embryological development provides evidence of phylogenetic relationships.

Fossil Evidence and Limitations

  • Fossils show similarities between organisms today and those now extinct.

  • Fossil record: incomplete record. It's biased toward species that existed a long time, were abundant, and had hard shells or skeletons.

Earth's Age and Radioactive Dating

  • Earth is estimated to be over 4.5 billion years old.

  • Rocks and sediments are dated through radioactive dating using isotopes.

Radioactive Isotopes and Half-Life

  • Isotopes: Atoms of the same element with different numbers of neutrons.

  • Radioactive isotopes undergo radioactive decay with a specific half-life.

  • Half-life: Time for half of an isotope sample to decay.

Carbon-14 Dating

  • Living things take in carbon-12 and carbon-14 constantly.

  • When dead, carbon uptake stops, and carbon-14 decays.

  • Limited to organic remains less than 75,000 years old.

Dating Older Rocks and Fossils

  • Use isotopes with longer half-lives (Potassium-40, Uranium-238).

  • Date rock layers by measuring radioactive isotopes.

  • Assign relative age to fossils found in those layers.