Chapter 20

Chapter 20: Communities

Part 1: Learning Outcomes

  1. Basic Definition of Community
    • Understand the term "community" in ecological context.
  2. Types of Interactions within Communities
    • Familiarize with three major interaction types: competition, symbiosis, and predation.
  3. Niche Definition
    • Grasp the concept of a niche and the impact of competition on species' niches.
  4. Coevolution
    • Understand coevolution and its role in symbiotic relationships.
  5. Predator/Prey Relationships
    • Learn about predator-prey interactions and coevolutionary dynamics.
  6. Ecological Succession
    • Recognize how succession contributes to biodiversity in communities.

Definitions

  • Population:
    • Refers to all members of a single species that live together in a specified geographic region.
  • Community:
    • Encompasses all species living in a particular geographic area.
  • Ecosystem:
    • The combination of a community and all its nonliving elements interacting with it.
  • Biosphere:
    • The global ecological system integrating all ecosystems.

Community Interactions

  • Interactions between populations within communities influence both ecological and evolutionary processes.
  • These interactions shape the community's structure and function.
  • Individualistic View:
    • Suggests that a community is merely a collection of species coexisting in one location.
  • Holistic View:
    • Views a community as a superorganism, with species functioning collectively as an integrated whole.
  • Assessing interactions within a community can be complex; biologists prioritize measurable interactions.
  • The concept that the whole is greater than the sum of its parts is fundamental in ecological studies.

Types of Community Interactions

  1. Competition:
    • Occurs when two or more populations utilize the same limited resource.
  2. Symbiosis:
    • Involves two or more species existing in close association, forming various types of relationships.
  3. Predation:
    • Describes the process where members of one species consume those of another species.

Competition

Niche vs. Habitat
  • Habitat:
    • The physical location where organisms reside.
  • Niche:
    • Represents all the ways an organism utilizes its habitat; it describes the organism's role within its environment.
  • Similar organisms often compete for resources in a habitat but can coexist by dividing their niches.
Fundamental vs. Realized Niche
  • Fundamental Niche:
    • The total niche an organism can potentially occupy.
  • Realized Niche:
    • The specific niche an organism occupies in the presence of competitors.
Competitive Dynamics
  • Gause's Principle:
    • States that no two species can occupy the same niche indefinitely under competition for limited resources.
    • Coexistence requires niche differentiation in one or more attributes. Lack of differentiation may lead to competitive exclusion, causing one species to become locally extinct.
Gause's Experiments
  • Conducted experiments with three Paramecium species to demonstrate competitive exclusion and niche differentiation.
  • When grown alone, all species flourished, but together, P. aurelia outcompeted P. caudatum. P. caudatum and P. bursaria coexisted by adjusting their realized niches.
Resource Partitioning
  • Describes how similar species avoid competition through utilizing resources differently (changing their realized niches).
  • Sympatric Species:
    • Closely related species in the same geographical area, avoiding competition through resource partitioning.
  • Allopatric Species:
    • Closely related species that avoid competition by existing in different geographical areas.
Case Studies
  • Example of Anolis lizards in the Caribbean partitioning resources based on microhabitat use.
  • Study of Geospiza finches on the Galapagos Islands showing phenotypic differences in beak size as a method of reducing competition for food.

Symbiosis

Coevolution
  • Describes the evolutionary adaptations of two or more species in response to one another.
  • Three primary forms of symbiosis include:
    1. Mutualism:
      • Both species benefit from the relationship.
    2. Commensalism:
      • One species benefits while the other is neither helped nor harmed.
    3. Parasitism:
      • One species benefits at the expense of the other.

Predation

Relationships
  • Predator:
    • The organism that consumes another organism (the prey).
  • Prey:
    • The organism that is consumed by the predator.
Predator-Prey Cycles
  • Predator and prey populations exhibit cyclical dynamics, often depicted graphically (e.g., snowshoe hare and lynx populations over time).
Coevolution in Predatory Relationships
  • When predator and prey are mutually dependent (often through an arms race dynamic), they influence each other's evolution.
  • Defensive adaptations:
    • Plants may develop thorns or toxic chemicals, while animals may evolve coloration, mimicry, or behavioral strategies.
Types of Mimicry
  1. Müllerian Mimicry:
    • Two harmful species resemble each other, enhancing predator avoidance.
  2. Batesian Mimicry:
    • A harmless species imitates a harmful species to deter predators.
  3. Self Mimicry:
    • An organism's body part resembles another part within the same organism, aiding in deception or defense.

Keystone Predators

  • Defined as predators crucial for maintaining species diversity in an ecosystem by controlling the populations of competitively dominant species.

Ecological Succession

  • Refers to a sequential process of community member replacement leading to a stable final ecological state.
  • Primary Succession:
    • Occurs in lifeless areas where soil has yet to form.
  • Secondary Succession:
    • Takes place in areas where disturbances have occurred but soil and some organisms remain.
Pioneer Species
  • Initial colonizers that modify the environment, facilitating further succession toward a climax community.
Disturbances
  • Natural events may reduce dominant populations, allowing for new communal dynamics.
Invasive Species
  • Non-native species that can hinder native biodiversity and disrupt successional processes.
  • Restoration efforts often focus on controlling invasives to promote native plant recolonization and habitats.
Ongoing Succession
  • The progression of ecological succession continues, occasionally reverting to earlier stages due to further disturbances, reinforcing the dynamic nature of ecosystems.