Chapter 20
Chapter 20: Communities
Part 1: Learning Outcomes
- Basic Definition of Community
- Understand the term "community" in ecological context.
- Types of Interactions within Communities
- Familiarize with three major interaction types: competition, symbiosis, and predation.
- Niche Definition
- Grasp the concept of a niche and the impact of competition on species' niches.
- Coevolution
- Understand coevolution and its role in symbiotic relationships.
- Predator/Prey Relationships
- Learn about predator-prey interactions and coevolutionary dynamics.
- Ecological Succession
- Recognize how succession contributes to biodiversity in communities.
Definitions
- Population:
- Refers to all members of a single species that live together in a specified geographic region.
- Community:
- Encompasses all species living in a particular geographic area.
- Ecosystem:
- The combination of a community and all its nonliving elements interacting with it.
- Biosphere:
- The global ecological system integrating all ecosystems.
Community Interactions
- Interactions between populations within communities influence both ecological and evolutionary processes.
- These interactions shape the community's structure and function.
- Individualistic View:
- Suggests that a community is merely a collection of species coexisting in one location.
- Holistic View:
- Views a community as a superorganism, with species functioning collectively as an integrated whole.
- Assessing interactions within a community can be complex; biologists prioritize measurable interactions.
- The concept that the whole is greater than the sum of its parts is fundamental in ecological studies.
Types of Community Interactions
- Competition:
- Occurs when two or more populations utilize the same limited resource.
- Symbiosis:
- Involves two or more species existing in close association, forming various types of relationships.
- Predation:
- Describes the process where members of one species consume those of another species.
Competition
Niche vs. Habitat
- Habitat:
- The physical location where organisms reside.
- Niche:
- Represents all the ways an organism utilizes its habitat; it describes the organism's role within its environment.
- Similar organisms often compete for resources in a habitat but can coexist by dividing their niches.
Fundamental vs. Realized Niche
- Fundamental Niche:
- The total niche an organism can potentially occupy.
- Realized Niche:
- The specific niche an organism occupies in the presence of competitors.
Competitive Dynamics
- Gause's Principle:
- States that no two species can occupy the same niche indefinitely under competition for limited resources.
- Coexistence requires niche differentiation in one or more attributes. Lack of differentiation may lead to competitive exclusion, causing one species to become locally extinct.
Gause's Experiments
- Conducted experiments with three Paramecium species to demonstrate competitive exclusion and niche differentiation.
- When grown alone, all species flourished, but together, P. aurelia outcompeted P. caudatum. P. caudatum and P. bursaria coexisted by adjusting their realized niches.
Resource Partitioning
- Describes how similar species avoid competition through utilizing resources differently (changing their realized niches).
- Sympatric Species:
- Closely related species in the same geographical area, avoiding competition through resource partitioning.
- Allopatric Species:
- Closely related species that avoid competition by existing in different geographical areas.
Case Studies
- Example of Anolis lizards in the Caribbean partitioning resources based on microhabitat use.
- Study of Geospiza finches on the Galapagos Islands showing phenotypic differences in beak size as a method of reducing competition for food.
Symbiosis
Coevolution
- Describes the evolutionary adaptations of two or more species in response to one another.
- Three primary forms of symbiosis include:
- Mutualism:
- Both species benefit from the relationship.
- Commensalism:
- One species benefits while the other is neither helped nor harmed.
- Parasitism:
- One species benefits at the expense of the other.
- Mutualism:
Predation
Relationships
- Predator:
- The organism that consumes another organism (the prey).
- Prey:
- The organism that is consumed by the predator.
Predator-Prey Cycles
- Predator and prey populations exhibit cyclical dynamics, often depicted graphically (e.g., snowshoe hare and lynx populations over time).
Coevolution in Predatory Relationships
- When predator and prey are mutually dependent (often through an arms race dynamic), they influence each other's evolution.
- Defensive adaptations:
- Plants may develop thorns or toxic chemicals, while animals may evolve coloration, mimicry, or behavioral strategies.
Types of Mimicry
- Müllerian Mimicry:
- Two harmful species resemble each other, enhancing predator avoidance.
- Batesian Mimicry:
- A harmless species imitates a harmful species to deter predators.
- Self Mimicry:
- An organism's body part resembles another part within the same organism, aiding in deception or defense.
Keystone Predators
- Defined as predators crucial for maintaining species diversity in an ecosystem by controlling the populations of competitively dominant species.
Ecological Succession
- Refers to a sequential process of community member replacement leading to a stable final ecological state.
- Primary Succession:
- Occurs in lifeless areas where soil has yet to form.
- Secondary Succession:
- Takes place in areas where disturbances have occurred but soil and some organisms remain.
Pioneer Species
- Initial colonizers that modify the environment, facilitating further succession toward a climax community.
Disturbances
- Natural events may reduce dominant populations, allowing for new communal dynamics.
Invasive Species
- Non-native species that can hinder native biodiversity and disrupt successional processes.
- Restoration efforts often focus on controlling invasives to promote native plant recolonization and habitats.
Ongoing Succession
- The progression of ecological succession continues, occasionally reverting to earlier stages due to further disturbances, reinforcing the dynamic nature of ecosystems.