Fossils, Radiometric Dating, and the Origin of Life
Comparative Biology and the Fossil Record
Reconstructing evolutionary history through living organisms provides cell, morphological, and molecular data, but lacks extinct species, time dimension, and environmental context.
The fossil record supplies the time context, extinct lineages, and environmental background essential to complete phylogenetic hypotheses.
Radiometric Dating
Carbon dating measures the ratio of radioactive Carbon-14 () decaying into stable Nitrogen-14 ().
The half-life of Carbon-14 is .
Carbon dating is reliable only for samples younger than approximately .
Requirements for Abiogenesis
Earth formed approximately , with life originating around to .
Four core conditions were required for chemical evolution:
Little or no free molecular oxygen ().
Energy sources (lightning, volcanic activity, cosmic/UV radiation).
Chemical building blocks (, , , , , , ; biogenic elements CHONPS).
Sufficient time.
The earliest organisms were prokaryotic and anaerobic, traceable to a shared lineage known as the Last Universal Common Ancestor (LUCA).
Hypotheses on the Origin of Life
Primordial Soup Hypothesis & Miller-Urey Experiment (1953):
A. Oparin and J. Haldane proposed organic molecules could form from inorganic raw materials.
Miller and Urey demonstrated that circulating gases (, , , ) exposed to electrical sparks without oxygen produce amino acids, sugars, and organic building blocks.

Clay Surface / Primordial Pizza Hypothesis:
Clay minerals in volcanic rocks bind and stabilize nucleotides on surfaces, concentrating them close enough to form small polymers.
Iron-Sulfur World Hypothesis:
Proposed by Günter Wächtershäuser (1992).
Life originated near deep-sea hydrothermal vents (black smokers and alkaline vents), where iron sulfide minerals acted as catalysts for autocatalytic carbon fixation.