Study Notes on Cladistics and Phylogeny
Class Schedule and Tests
Final Test Schedule
The last actual test of the year is scheduled for April 21.
A common misconception among other classes is that the algae test was the last one, which is incorrect.
After the final test, there will be a major project, the details of which will be explained later.
Phylogenetics and Cladograms
Objective of Phylogeny
Phylogeny involves determining evolutionary relationships among organisms.
The aim is to classify organisms in a way that relates to their evolutionary lineage.
Definition of Cladograms
Cladograms are diagrams that illustrate these evolutionary relationships.
They are key tools in depicting how groups of organisms are linked through common ancestry.
Historical Context of Cladistics
Initial Methods for Classifying Organisms
Early classification relied on physical characteristics due to the lack of technological means to study genetic information.
Over time, understanding of evolutionary relationships has shifted from physical traits to genetic data.
Evolution of Thoughts on Traits
Physical traits can change over time, complicating classification efforts.
For example, dolphins were once classified with fish, but are distinct due to evolutionary differences.
Cladistics Terminology
Clade
Definition: A clade is a group that includes an ancestor and all of its descendants.
Clades can represent single branches or multiple branches on a cladogram.
Nodes
Nodes are points on a cladogram where two organisms or species diverge.
Each node represents a speciation event in evolutionary history.
Traits and Their Development in Cladistics
Derived Traits
Definition: Derived traits are evolutionary innovations that appear as organisms develop over time.
These traits are crucial for constructing cladograms, as each derived trait indicates shared characteristics among groups.
Example:
In the discussion of legs, it was noted that all organisms in a certain clade must possess the trait described (e.g., legs).
However, exceptions, such as snakes (which have lost legs), need to be annotated with explanations, such as “loss of legs.”
Cartilage and Bone in Evolution
Cartilage
Definition: Cartilage is a flexible tissue that serves as a precursor to bone in many species.
Characteristics: It is easier to produce, more flexible, and offers decent strength.
It plays a role in the early development of creatures, including humans, which begin life with cartilage before bone formation.
Bone
Bones are strong, rigid structures that evolved from ancestral cartilage.
Interpreting Cladograms
Outgroup and Ingroup
Outgroup: The least closely related group in a cladogram, sometimes used as a common ancestor.
Ingroup: The primary focus group in the cladogram; includes the organisms of interest among the clades.
Cladogram Orientation
The arrangement of cladograms can vary without altering the relationships depicted.
Similar relationships can be represented in different orientations, like trees.
Building Cladograms Today
Modern Techniques
Genetic data now plays a critical role in updating and correcting ancestral relationships.
Morphological evidence is often supplemented or modified by genetic evidence.
Example: Birds and reptiles were classified apart traditionally, but modern genetics shows they are more closely related than previously thought.
Principle of Parsimony
Definition: The principle of parsimony dictates that the most credible cladogram is the one with the fewest changes.
This principle ensures that traits should not appear and disappear randomly across branches.
Consistent presence of traits among branches reinforces the credibility of the cladogram.
Practical Assignments
Dichotomy Assignment
Students will complete a dichotomy task focusing on identifying the scientific names of various organisms.