Population Ecology

Introduction to Population Ecology
  • Population ecology studies how biotic (living) and abiotic (non-living) environmental factors influence population abundance, dispersion, and age structure.
  • Abiotic factors, like temperature, are non-living environmental components that affect populations.
Characteristics of Populations
Definition of a Population
  • A population is a group of individuals of a single species in a specific geographic area, characterized by its population size (NN).
Density
  • Density is the number of individuals per unit area or volume. It changes due to births, deaths, immigration (influx), and emigration (outflux).
Dispersion
  • Dispersion is the spatial pattern of individuals in a population.
  • Three types are recognized:
    • Clumped: Individuals gathered in patches, often due to resource distribution or social behavior.
    • Uniform: Evenly spaced individuals, often due to territoriality or competition.
    • Random: Individuals positioned independently, occurring in the absence of strong attractions or repulsions or in uniform environments.
Demography
  • Demography is the study of population vital statistics (birth rates, death rates, migration rates) and their changes over time.
Life Tables
  • Life tables track survival and reproductive rates of a cohort (same-aged group) through its life.
Survivorship Curves
  • Survivorship curves graph the proportion of a cohort surviving at each age, categorized into three types:
    • Type I: Low early/middle death rates, high older death rates (e.g., large mammals, humans).
    • Type II: Constant death rate throughout life (e.g., some rodents).
    • Type III: High death rates for the young, lower for survivors (e.g., oysters, many fish).
Estimating Population Size
  • Population size is estimated using sampling techniques like:
    • Extrapolation from small samples.
    • Index of population size (e.g., nests, tracks).
    • Mark-Recapture Method: For mobile species, where N=snxN = \frac{sn}{x} (N = estimated population size, s = marked in first sample, n = total in second sample, x = marked in second sample).
Population Growth Models
  • Population growth patterns are shaped by biotic and abiotic factors.
Exponential Growth
  • Occurs under ideal conditions (unlimited resources), leading to a J-shaped growth curve.
  • The instantaneous rate of change is described by dNdt=rN\frac{dN}{dt} = rN, where rr is the instantaneous per capita growth rate and NN is the current population size.
Logistic Growth
  • Accounts for limited resources; populations cannot grow exponentially forever.
  • Carrying Capacity (KK): The maximum population size an environment can sustain.
  • Results in an S-shaped growth curve; growth slows as NN approaches KK.
  • Formula: dNdt=rN(KN)K\frac{dN}{dt} = rN\frac{(K-N)}{K}, where KK is carrying capacity.
  • When N=KN = K, population growth stops (dNdt=0\frac{dN}{dt} = 0).
  • Comparison: Exponential growth assumes unlimited resources (J-curve); Logistic growth accounts for limits and carrying capacity (S-curve).
Population Regulation
Density-Dependent Factors
  • Birth rates fall and death rates rise with increasing population density, acting as negative feedback.
  • Examples: Competition for resources, disease spread, predation, territoriality, toxic wastes, and intrinsic physiological factors.
Density-Independent Factors
  • Birth and death rates are unaffected by population density.
  • Examples: Natural disasters like floods, droughts, and wildfires.
Importance for Population Size Regulation
  • Only density-dependent factors regulate population size, causing it to stabilize or decrease if thresholds are exceeded.
Life Histories
Definition
  • An organism's life history comprises traits characterizing its reproduction and survival schedule, shaped by natural selection.
Key Components of Life History
  • Critical components include:
    1. Age at first reproduction.
    2. Frequency of reproduction.
    3. Number of offspring per reproductive episode.
Reproductive Strategies
  • Semelparity: Organisms reproduce once in their lifetime and then die.
  • Iteroparity: Organisms reproduce multiple times throughout their lifespan.