Social Perception and Rage and Agression
Lecture 18
Processing facial identity
FFA (fusiform ventral gyrus) in ventral occipitotemporal cortex
Part of identity- things that dont change
Processing facial expression
STS superior temporal sulcus in temporal lobe
Feeds to amygdala to decide if there is a threat
Responds to any type of biological motion
In monkeys:
STS (expression) and area IT (identity)
Difference in humans and monkeys have to do with visual cortex areas
Have extra cortex that has pushed visual areas like area IT a nd displaced it underneath the brain
Expansion of temporal lobe due to language
Facial symmetry
Symmetry may indicate phenotypic quality and how well an individuals genome can resist disease and maintain normal development in the face of environmental perturbation
If soeone is attrcated to symmetric face theta re attracted to underlining genetic quality of the person with the symmetric fae
Testosterone and estrogen during puberty affect the face anatomy
Health and attractiveness
Mesolimbic DA and attracitiveness
When people see attractive person it activates mesolimbic DA reward
NA and MOFC , makers attractivenessa rewsarding
Lecture 19 rage and aggression
Testosterone - motivates the pursuit and defense of btoh mates and social status
Only facilitates aggression when aggression is a path to sical status
Often related to competition for social status
Female reproductive development
Gonads develop into ovaries
Mullerian ducts develop
Fallopian tube
Uterus
Cerix
Wolffian ducts regress
Prenatal ovarian hormones are not involved in this process
External reproductive system is also differentiating:
Clit, labia, vagina
Male reproductive development
At about 6 weeks, TDF (testes determination factor) gene on Y chromosome causes gonads to develop into testes
Testes produce etstosterone starting at about 8 weeks, peaking at 14 weeks
Levels then decline until second peak 2 months after birth
Low levels until puberty
Wolfian ducts develop
Vas deference
Seminal vessicle
Prostate gland
MIH from testes causes regression of mullerian ducts
Process of masculinizing internal reproduction system structure is dependent of gonadal hormones
External genitalia - dependent on dihydrotestosterone
Penis and scrotum
Steroid hormones makes neurons grow
Testosterone and estrogen promote neuritic growth in cultured hypothalamic neurons from newborn mice
Changes anatomy of the brain
SDN POA in rats
The sexually dimorphic nucleus of the preoptic area of the hypothalmus (SND-POA) is five times larger in male vs female rats
Organizational effects of T
Give female rates testosterone, will develop male sized SDNPOA
Castrate male rates (no more T) will develop female sized SDN POA
Mouse aggression depends on both organizational and activational effects of testosterone
Female mice show male levels of aggression if (have to do both)
Exposed to testosterone on first day of life (organizational effects/ changes brain anatomy)
Given testosterone as adults (activational effects)
Testes acts later in life that presumably acts in neural circuitry that was organized earlier by testosterone (activating what was already there)
Organizational effects of T on aggression
Male monkeys show more rough and tumble play as juveniles
Exposing female fetuses to testos in utero increases rates of rough and tumble play
Blocking androgen (what testos binds to) receptors in male fetuses in utero decreases rough and tumble play
If woman was having trouble with pregnancy dr will prescribe progesterone which maintains the lining of uterus
Progestorone also binds to and activaets androgen receptor (acts like testos in utero)
Babies developing would be exposed to testos
These babies had a higher score of aggression when compared to unexposed siblings
Activational effects of T on aggression
During mating season, rhesus macaques show
Increased testes size
Increased plasma testos concentrations
Increases muscle mass
Increased ratesof aggression
Association between high levels of aggression and testos
Relationship between T and aggression is bidirectional
Physiological injections of T exaggerate existing patterns of aggression
Promssive effects
Aggression can affect T
Aggression can cause increase in T
Winning vs losing
Activation effects of T on human aggression
As levels of T in adulthood increase aggressive behavior in adulthood
Higher t in violent vs nonviolent male and female cirminals
Correlates with response to provations
Ultimate game - when you choose to reject, you re punishing other person (aggression)
Linked testos to decisions made in game
Rejected - higher T levels
Some studies fail to find associations between t levels and aggression in humans
Might be bcecause T is more about motivating people to compete for social status. Path for status doesnt really come from aggression in humans
About maintenance and pursuit of status than aggression. For animals the way to get status is through aggression
Overall, a weak but significant association
T may be more directly related to the maintenance and pursuit of social status
Neurological effects of T
Cells that bind to T in monkeys are concentrated in: (regions gave a lot of cells that have androgen receptors
Midbrain (PAG)
Hypothalamus
Amygdala
Defensive rage circuit in the domestic cat
Electrical stimulation of midbrain (PAG), medial hypothalamus, or maygdala produce affective aggressive response
Growling, hissings, piloerection
Back arch
Paw strike
Stria terminalis links amygdala and hypothalamus
T increases firing frequency if it is already firing
Stria - white matter pathway where bunch of different neurons send their axons down with same connection
Ation potentials travel down those axons during rage response
T and the amygdala response to threatening faces
If have higher T, have stronger amygdala response
The more active the orbital frontal cortex was, the less likely they were to reject unfair offer and have aggressively
At the medial frontal cortex, T inhibits it (releases control of MFC and amygdala) and get more aggression)
MFC helps restrain the amygdala and regulate it (inhibitory effect on amygdala
Serotonin
There are nuclei of serotonin neurons that send axons throughout the brain and release serotonin in manu different parts of the brain
Called raf a nuclei
Serotonin and aggression in free ranging rhesus macaques
Capture monkeys and take sample of cerebral spinal fluid sample which tells you about the brain
Within sample of spinal fluid can measure metabolite of serotonin
Low CSF 5 HIAA (metabolite of brain serotonin) is associated with
Severe unprovoked aggression
Deficits in impulse controls
Serotonin has imposing effects to T
High T / lwo 5 HIAA associated with highest rates of aggression
T provides competitive drive
Serotonin regulates the threshold for aggression
Testos provided competitive drive/mtoivation and serotonin regulated the threshold for aggression
Low serotonin, low threshold to respond aggressively
Neural circuitry of emotion regulation
Men with damage to OFc are more likely than men with damage in other brain regions to have outburst of rage at family members, friends, and colleagues (lost impulse control)
Serotonin agonists increase glucose metabolism in OFC
This increase is blunted in patients with aggressive impulsive personaility disorders
There are lots of serotonin receptors in OFC may act there to inhibit aggression
Serotonin might act opposite of T, control limbic system response
MAOA allele and aggression
Monoamine oxidase enzyme
Clears away excess NT from synapse, breaks down monoamine NT (DA, epinephrine, serotonin)
Evidence:
Large dutch kindred with MAOA mutation with persistent and extreme reactive aggression in some males
Male MAOA knockout mice are hyperaggressive
Men with L allele of MAOA are more prone to impulsive violence, particularly if they were abused as children
Exampels of egen by environment interaction - gene more likely to have an affect when combined with certain aharacteric of environment
OT and maternal aggression
We see mothers aggressively defending offspring
OT facilitates her not being overcome by fear and anxiety so she can face predator (lots of OT from birth)
OT decreases firing rates of neurons in amygdala (which has anxiolatic/ relaxing effects)