Evolution of Plant Diversity Notes
Evolution of Plant Diversity
- Common Ancestor: Plants and green algae (charophytes) evolved from a shared ancestor; both are multicellular and photosynthetic eukaryotes.
- Examples:
- Chara: A complex charophyte
- Coleochaete: A simpler charophyte
Key Traits of Plants
- Five key traits appear in nearly all land plants but are absent in charophytes:
- Alternation of Generations: Life cycle alternates between gametophyte (haploid) and sporophyte (diploid) stages.
- Multicellular, Dependent Embryos: The embryonic stage develops within the female gametophyte.
- Walled Spores in Sporangia: Spores are produced in protective structures called sporangia.
- Multicellular Gametangia: Structures that produce gametes are multicellular.
- Apical Meristems: Regions of actively dividing cells at the tips of roots and shoots for growth.
- Additional Derived Traits:
- Cuticle: A waxy layer that reduces water loss.
- Stomata: Pores for gas exchange, typically on the leaves.
- Mycorrhizae: Fungal associations that enhance nutrient uptake.
Plant Life Cycle Overview
- Contains alternating haploid (n) and diploid (2n) generations, involving both mitosis and meiosis.
- Gametophyte (n): Produces gametes through mitosis.
- Sporophyte (2n): Develops from the zygote after fertilization; produces spores via meiosis.
Adaptations to Life on Land
- Moisture Retention: Mechanisms to maintain moisture within cells.
- Structural Support: Need for support in a nonbuoyant medium.
- Reproduction: Adaptations for reproduction without water.
- Anchoring: Roots anchor the plant, absorbing nutrients and water from the soil.
Comparison: Aquatic vs. Terrestrial Adaptations
- Vascular Tissue: Present in terrestrial plants for better nutrient and water transport.
- Roots: Essential for anchoring and moisture absorption.
- Leaves: Increase surface area for photosynthesis.
- Stomata: Allow gas exchange while minimizing water loss.
Key Events in Plant Evolution
- Origin of Land Plants (about 470 mya): Nonvascular plants (bryophytes) such as mosses, hornworts, and liverworts.
- Origin of Vascular Plants (about 425 mya): Seedless vascular plants like ferns with flagellated sperm.
- Origin of Seed Plants (about 360 mya): Development of vascular tissues with lignin providing structural support, including gymnosperms and angiosperms.
Bryophytes: Nonvascular Plants
- Three Phyla:
- Liverworts (Hepatophyta)
- Mosses (Bryophyta)
- Hornworts (Anthocerophyta)
- Dominated by the gametophyte generation, which is larger and longer-living than the sporophyte.
Seedless Vascular Plants
- Lycophytes and Monilophytes:
- Examples:
- Club Mosses (Lycophyta)
- Ferns (Monilophyta)
- Adaptations:
- Lignin for increased structural support.
- Vascular tissues (xylem and phloem) for transport.
Introduction of Seed Plants
- Key Adaptations:
- Seeds: Protective structures surrounding the embryo and food supply.
- Pollen: Male gametophytes that can be dispersed without water, aiding in reproduction success.
- Types of Seed Plants:
- Gymnosperms: Produce seeds in cones.
- Angiosperms: Seeds develop in protective ovaries, leading to fruit formation.
Angiosperm Diversity
- Flower Structure:
- Flowers contain male and female reproductive structures.
- Serve as sites for pollination and fertilization.
- Life Cycle: Characterized by double fertilization, where one sperm fertilizes the egg while the other combines with polar nuclei to form endosperm.
- Fruit Development: Fruits aid in seed dispersal through various mechanisms: wind, animals, or edible fruit consumption.