Notes on DNA Extraction and Applications in Molecular Biology
INTRODUCTION
- Overview of DNA extraction process: removal of DNA from other cellular components.
STEP 1: LYSIS
- Definition: The process of breaking down the cell to release DNA from the nucleus.
- Alternatives: Known as cell lysis or cell disruption.
- Methods:
- Physical method:
- Targets cells with strong membranes like plant cells.
- Techniques include:
- Blender
- Mortar and pestle (ครกและสาก)
- Cutting (การตัด)
- Sonicator (คลื่นเสียงความถี่สูง)
- Chemical method:
- Destroys nuclear membrane (nuclear membrane lysis).
- Uses detergents, e.g.:
- Sodium dodecyl sulfate (SDS)
- Lauryl sulfate
- Employs enzymes to break bonds in the proteins, e.g.:
- Proteinase K
- Cellulase
- Lyticase
- Lysozyme
STEP 2: PRECIPITATION
- Definition: The process of separating DNA from cellular debris after lysis.
- Post-lysis content:
- Cell extract containing DNA + proteins + detergents + other chemicals.
- Mechanism:
- Sodium ions (Na⁺) neutralize the charges of DNA, stabilizing it.
- Alcohol (e.g., isopropanol or ethanol) causes DNA to precipitate as it is insoluble in alcohol.
- Proteases digest proteins that are bound to DNA.
- Filtration aids in separating contaminants.
STEP 3: WASH / PURIFICATION
- Process: The precipitated DNA is washed with cold alcohol.
- Centrifugation: Used to remove contaminants such as:
STEP 4: RESUSPENSION
- Definition: The step where DNA pellet is dissolved in a buffer solution.
- Common buffers used:
- Tris buffer
- TE buffer
- Double distilled water
- Outcome: DNA is ready for subsequent applications.
1. UV ABSORBANCE
- Method: Measures absorbance of DNA at specific wavelengths.
- Wavelengths:
- DNA and RNA absorb at 260 nm.
- Proteins absorb at 280 nm.
- Quality indicators:
- Ratio A260/A280 to determine purity of DNA (ideal = 1.8–2.0).
- Ratio A260/A230 indicates contamination (ideal > 1.5).
- Advantages:
- Quick and does not involve chemicals.
- Limitations:
- Low sensitivity when DNA concentrations are low.
- Cannot separate DNA from RNA.
2. FLUORESCENT DYES
- Function: Used to bind specifically to double-stranded DNA (dsDNA) with examples like:
- Features:
- Higher sensitivity compared to UV absorbance.
- Requires a standard curve for quantification.
3. AGAROSE GEL ELECTROPHORESIS
- Purpose: Evaluates quantity and integrity of DNA across various sizes.
- Process:
- DNA is run on an agarose gel stained with an intercalating dye such as ethidium bromide.
- Analysis:
- Band intensity compared against a DNA ladder.
- Benefits:
- Visualizes DNA size and integrity; detects contamination.
- Limitations:
- Cannot quantify protein or salt contamination.
4. CAPILLARY ELECTROPHORESIS
- Overview: Similar to agarose gel electrophoresis but in an automated system.
- Requirements:
- Only 1–2 µL of sample used.
- Uses fluorescent dyes for visualization.
5. DIPHENYLAMINE METHOD
- Reaction: Diphenylamine reacts with deoxyribose under acidic conditions.
- Result: Produces a blue compound measurable at 595 nm.
- Limitations:
- Low sensitivity, time-consuming, and not commonly used now.
6. qPCR / RT-PCR
- Function: Quantifies DNA during amplification using fluorescence.
- Standard curve: Necessary for accurate quantification.
- Advantages:
- High sensitivity and accuracy.
SUMMARY
- DNA extraction is a fundamental process in medical, biomedical science, and biotechnology.
- Main Steps: lysis → precipitation → purification → elution.
- Method selection depends on sample type, age, and amount.
APPLICATIONS OF MOLECULAR BIOLOGY TECHNIQUES IN MEDICINE
A. GENETIC DISEASE AND DISEASE IDENTIFICATION
- Significance of molecular genetic testing:
- Early detection
- Predictive testing
- Carrier screening
- Personalized medicine
- Disease prognosis.
- Methods used:
- Targeted testing
- Gene panel
- Whole-exome sequencing (WES)
- Whole-genome sequencing (WGS)
- Chromosomal tests (e.g., FISH, CMA, karyotype)
- Gene expression tests
- Biochemical tests.
B. FORENSICS
- Applications include:
- Paternity tests
- Victim/suspect identification.
C. PRENATAL SCREENING AND GENETIC COUNSELING
- Focus: Genetic mutation identification, carrier screening, family history analysis.
- Typical methods:
- Non-invasive prenatal testing (NIPT)
- Chorionic villus sampling (CVS)
- Amniocentesis.
- Example: Mycobacterium tuberculosis.
- Processes involved:
- Sample collection
- DNA extraction
- PCR amplification
- Interpretation of results.
E. PHARMACOGENOMICS
- Study of genetic variation affecting drug response leading to:
- Personalized medicine
- Gene expression analysis
- Gene editing techniques such as CRISPR-Cas9
- Next Generation Sequencing (NGS).
F. CANCER RESEARCH AND DIAGNOSTICS
- Focus areas:
- Cancer biology
- Genomic profiling
- Liquid biopsy
- Imaging techniques (CT, MRI, PET)
- Precision medicine.
- Future trends to watch:
- Immunotherapy
- Targeted therapy
- Artificial Intelligence (AI) in diagnostics
- CRISPR/Cas9 for gene editing and treatment.