Forest Ecology Notes
Forest Ecology
Disturbance
- A relatively event in and that alters the of , , and and causes changes in and/or the ___ ___.
Factors Influencing Disturbance Impact
- The impact of disturbance on an ecosystem depends on several factors:
- Size
- Pattern
- Severity
- Frequency
- Timing
- Intensity
- Type of disturbance
- These factors together constitute a ____.
Timing
- Timing determines impact.
- Example:
- Fire or freeze during bud break causes much more mortality.
- Fire in summer is more severe than in winter.
Disturbance Interactions
- The effect of one disturbance on the probability of another disturbance occurring (Picket and White 1985).
- Example:
- Beetle-killed spots within a loblolly pine stand may be relatively small in proportion to the stand.
- However, they are areas of high fuel loading, increasing the risk that a lightening-ignited wildfire will start in these areas and spread to the surrounding stand.
Perturbation and Response
- Perturbation: An external force that displaces a system from equilibrium.
- Response: Describes the direction and magnitude of change in system after perturbation.
- Resistance: Tendency of the system to remain in a reference state in the face of perturbation (i.e., maintain certain structures and functions despite being disturbed).
Recovery and Resilience
- Recovery: The extent to which a system returns to its original state after perturbation.
- Recovery depends on the magnitude of response and time since perturbation.
- Resilience: Rate at which a system returns to a reference state.
System Stability
- Systems that maintain properties despite disturbance (i.e., resistant) and return rapidly to their original state (i.e., resilient) are more stable and predictable.
State Changes
- Savanna state
- Woodland state
- Forest state
- TRIGGER: cause of change
- TIPPING POINT: threshold beyond which the system changes state
- ECOLOGICAL CONSEQUENCES
Disturbance and Community Equilibrium
- Disturbance maintains communities in a "non-equilibrium state" (never reaching equilibrium).
- By renewing colonizable space, disturbance allows the persistence of species that might otherwise go extinct due to competitive exclusion.
- Attribution: Joe Connell
Intermediate Disturbance Hypothesis
- Species diversity is low at low disturbance frequency because of competitive exclusion.
- Species diversity is higher at intermediate disturbance frequency due to a mix of good colonizer and good competitor species.
- Species diversity is low at high disturbance frequency because only good colonizers or highly tolerant species can persist.
Succession
- A directional change in ecosystem structure and function resulting from biotically-driven changes in resource availability following the formation of barren land (primary) or a disturbance (secondary).
- Organisms respond to changes in resources, and the presence of those organisms alters resources, creating a cascade effect.
Seral Stages
- Different stages of communities are seral stages that are distinct.
- Transitions between seral stages are usually gradual rather than abrupt.
- Each stage transforms the ecosystem’s environment, creating conditions that foster the next stage’s development.
Primary Succession
- Occurs when a community develops on a site without established biota (no soil, plants, or seeds).
Primary Succession (Glacier Example)
- Bare rock left after the retreat of a glacier.
- In time, mosses and lichens start to colonize the rock.
- As they die, organic matter is added to weathered rock particles, making simple soils.
- As the soils develop, grasses and small herbaceous plants start to grow.
- More organic matter is added, and roots of plants aid breakup of rock material.
- Deeper soils hold more water; small shrubs colonize these better soils.
- Nutrient availability increases
- More root action
- Eventually, trees establish, leading to the development of a climax community on mature soils.
Glacier Bay Succession
- Exposed rock
- Pioneer species: herbal rose and willow
- Alder bush
- Spruce
- Spruce-hemlock
- Moss, herbaceous plants
- Ponds and bogs
Soil Development During Primary Succession
- The depth of all major soil layers increased during the first 200 years of succession.
Species Succession
- Alder
- Willow
- Spruce
- Cottonwood
- Hemlock