Formation of Earth
Earth's formation took place approximately 4.6 billion years ago.
Early Earth experienced heavy bombardment by rocks, which vaporized water, delaying the formation of seas until around 4 billion years ago.
Early Atmosphere
The atmosphere at the time contained little oxygen and included
Water vapor
Volcanic gases such as nitrogen, carbon dioxide, methane, ammonia, and hydrogen.
Origin of Simple Cells
Hypothesized chemical and physical processes may have produced simple cells through a sequence of stages:
Abiotic synthesis of small organic molecules.
Joining of these small molecules into macromolecules.
Packaging of molecules into protocells.
Development of self-replicating molecules.
Key Figures and Experiments
Oparin & Haldane (1920s): Proposed the early atmosphere was a reducing environment.
Miller & Urey (1953): Conducted experiments demonstrating the abiotic synthesis of organic molecules in a reducing atmosphere.
Meteorites
A significant source of organic molecules, notably the Murchison meteorite:
Weighed over 100 kg and contained 80+ amino acids, along with lipids, simple sugars, and nitrogenous bases.
Fell in Victoria, Australia in 1969 and was a documented event.
5 Mass Extinctions
Hadean Eon (4.6-4.0 BYA)
Formation of Earth
Intense meteorite bombardment
Formation of Earth's crust and early oceans
Minimal evidence of life; origin of simple organic molecules.
Archaean Eon (4.0-2.5 BYA)
Formation of the first solid continents
Emergence of prokaryotic microbes (bacteria and archaea)
Photosynthesis by cyanobacteria released oxygen.
Proterozoic Eon (2.5 billion-541 million years ago)
The Great Oxygenation Event resulted in significant oxygen buildup.
Appearance of eukaryotes, simple algae, and soft-bodied organisms.
Phanerozoic Eon (541 MYA-present)
The Cambrian Explosion marked rapid diversification of life.
Development of hard-shelled organisms, complex plants, and animals.
Evolution of fish, insects, reptiles, mammals, and flowering plants.
Multiple mass extinctions, including the one that ended the dinosaurs.
Rise of humans in the Holocene epoch.
Hierarchy of Taxonomic Groups
Ranges from broad to narrow:
Domain
Kingdom
Phylum
Class
Order
Family
Genus
Species
Maximum Parsimony and Maximum Likelihood
Maximum Parsimony: Posits that the simplest tree requiring the fewest evolutionary events is likely the true phylogeny.
Maximum Likelihood: Uses probability rules regarding DNA changes over time, inferring the evolutionary tree that reflects the most probable sequence of events.
Orthologous vs. Paralogous Genes
Orthologous Genes:
Found as a single copy in the genome and homologous across species, diverging post-speciation.
Paralogous Genes:
Result from gene duplication, found in multiple copies within the genome, diverging within the species.
Horizontal Gene Transfer
Concept triggering major genetic changes between organisms.
Pardomorphosis
Refers to accelerated reproductive development compared to somatic development, where sexually mature species retain juvenile features.
Classification Criteria
Shape
Cocci: Round shape
Bacilli: Rod shape
Spirilli: Spiral shape
Reproduction
Asexual reproduction via binary fission.
Genetic exchange through conjugation, transformation, and transduction.
Nutrition
Autotrophs: Produce their food (e.g., cyanobacteria via photosynthesis).
Heterotrophs: Consume organic material.
Chemotrophs: Obtain energy from inorganic compounds.
Oxygen Requirements
Obligate aerobes: Require oxygen.
Obligate anaerobes: Cannot survive in the presence of oxygen.
Facultative anaerobes: Can survive with or without oxygen.
Gram Staining:
A classification method for bacteria:
Gram-positive: Thick peptidoglycan walls (stains purple).
Gram-negative: Thinner peptidoglycan and an outer membrane (stains pink).
Reproductive Methods
Binary Fission: A fast, asexual process resulting in clones; occurs every 1-3 hours under optimal conditions.
Conjugation: DNA transfer between bacteria through a pilus.
Transformation: Uptake of DNA from the environment.
Transduction: DNA transfer between bacteria via a virus.
Mutation Rates
Low mutation rates can lead to rapid accumulation of mutations due to shorter generation times and large populations.
Plasmids: Small, independently replicating DNA rings; often carry genes for antibiotic resistance, important for bacterial evolution.
Applications:
Used in bioremediation to remove pollutants from environments.
Experiments utilizing prokaryotes have advanced DNA technology.
Bacteria can aid in producing natural plastics.
Endospores: Tough, metabolically inactive spores that can survive extreme conditions for centuries.
Fimbriae: Hair-like appendages allowing prokaryotes to adhere to substrates or other individuals.
Pili: Longer appendages that facilitate DNA exchange between bacteria, particularly during conjugation.
Symbiosis: Close ecological interactions between two species, typically including a larger host and a smaller symbiont.
Mutualism: Both organisms benefit.
Commensalism: One benefits without harming or helping the other significantly.
Parasitism: One organism (the parasite) harms the host.
Pathogens: Bacteria responsible for many diseases in humans.
Exotoxins: Secreted toxins affecting hosts even without the bacterial presence.
Endotoxins: Released only upon bacterial death, leading to disease symptoms.
Nutritional Diversity:
Protists exhibit both sexual and asexual reproduction.
Photoautotrophs: Contain chloroplasts.
Heterotrophs: Absorb organic molecules.
Mixotrophs: Combine traits from both nutritional modes.
Excavata: Unique cytoskeleton with feeding grooves.
Diplomonads: Reduced mitochondria, anaerobic pathways, two identical nuclei, multiple flagella.
Parabasalids: Mitochondrial remnants (hydrogenosomes), energy via anaerobic biochemistry.
Euglenozoans: Diverse clade with predatory and photosynthetic species, distinguished by spiral flagellar rods.
SAR:
Stramenopiles: Photosynthetic organisms with dual flagella; includes diatoms and brown algae.
Alveolates: Membrane-enclosed sacs (alveoli) under plasma membranes; includes dinoflagellates, apicomplexans, ciliates.
Rhizarians: Includes amoebas; distinguishes themselves with threadlike pseudopodia.
Archaeplastida: Comprises red algae, green algae, and plants.
Red Algae: Multicellular, reddish due to phycoerythrin.
Green Algae: Closely related to land plants; include charophytes and chlorophytes.
Unikonta: Animals, fungi, and some plants.
Amoebozoans: Have lobe- or tube-shaped pseudopodia; includes slime molds and entamoebas.
Endosymbiotic Theory:
Proposes that mitochondria and plastids originated from engulfed prokaryotes by early eukaryotic ancestors, aligning mitochondria with alpha proteobacteria and plastids with photosynthetic cyanobacteria.
Fungi Roles:
Decomposers: Break down nonliving organic material for nutrient absorption.
Parasitic: Absorb nutrients from living hosts.
Mutualistic: Benefit host plants; mycorrhizae enhance nutrient absorption.
Body Structures:
Common structures include multicellular filaments and single-celled yeasts.
Fungi develop hyphae networks for nutrient absorption, reinforced by chitin.
Hyphal Networks:
Form extensive mycelium enhancing feeding efficiency.
Plasmogamy and Karyogamy:
Plasmogamy: Fusion of cytoplasm from different mycelia.
Karyogamy: Fusion of nuclei produces diploid cells, enabling genetic variation.
Recognized phyla:
Chytrids
Zygomycetes
Glomeromycetes
Ascomycetes
Basidiomycetes
Lichens: A symbiotic relationship between fungi and photosynthetic microorganisms (green algae or cyanobacteria).
Ergotism: A fungal infection noted for dramatic symptoms linked to historical events.
Fungal Infections: General term is mycosis, including common ailments like ringworm and athlete's foot.
Practical Uses:
Fungi hold importance in antibiotic production and cancer research.