AP BIOLOGY NOTES

Standard Deviation (SD)

  • A measure of how spread out the data is from the mean

Lower SD:

  • Data is closer to the mean

  • Greater likelihood that the independent variable (IV) is causing the changing in the dependent variable (DV)

Higher SD:

  • Data is more spread out from the mean

  • Factors, other than IV, are influencing DV

Calculating SD:

  1. Calculate the mean

  2. Determine the difference between each data point and the mean

  3. Square the differences

  4. Sum of the squares

  5. Divide sample size (n) minus 1

  6. Take the square root

Standard Error (SE):

  • How well the mean of a sample estimates the true mean of population

  • Measure of accuracy, if true mean is known

  • Measure of precision, if true mean isn’t known

  • Accuracy: How close a measured value is to the actual (true) value

  • Precision: How close the measured values are to each other

  • Look for overlap of error lines

    • If they overlap, the difference isn’t significant

    • If they don’t overlap, the difference may be significant

Chi-Squared (X²) Analysis:

  • A test that measures how a model compares to actual observed data

  • In genetics, you can predict genotypes based on probability (expected results)

  • Chi-Squared is a form of statistical analysis used to compare the actual results (observed) with the expected

    • Never take square root of X²

  • If the expected and observed values are the same - X² = 0

  • By calculating the X² value, you determine if there is a statistically significant difference between the expected and actual values

Null Hypothesis:

  • Your null hypothesis states that there is no difference between the observed and expected values

  • You will either accept or reject the null based on the chi-squared value that you determine

  • Calculating

    • Determine what your expected and observed values are

    • Actual values should be something you get from the data - no calculations

    • Expected is based on probability

      • Make a table

Degrees of Freedom:

  • # of categories minus 1

  • If X² > critical value, there is a significant difference

Animal Behavior:

  • Based on physiological systems and processes

  • A behavior is the nervous system’s response to a stimulus

  • Helps an animal

    • Obtain food

    • Find sexual reproduction partner

    • Maintain homeostasis

  • Ethology - Study of animal behavior

  • Behavioral Ecology - Study of ecological and evolutionary basis for animal behavior

  • Integrates proximate/ultimate explanations for animal behavior

  • Proximate causation, or “how” explanations

    • Environmental stimuli

    • Genetic, physiological, and anatomical

  • Ultimate causation or “why”

    • Evolutionary significance

Fixed Action Pattern:

  • A FAP is a sequence of unlearned, innate behaviors that is unchangeable

  • Once initiated, carried to completion

  • Triggered by an external cue - sign stimulus

Taxis:

  • Environmental cues trigger movement in particular direction

  • Taxis automatic, oriented movement toward/away from a stimulus

Kinesis:

  • Simple change in activity or turning rate in response to a stimulus

CER:

Claim: A statement that answers your question

Evidence: Scientific data to support the claim

  • Data needs to be appropriate (pictures, graph, table)

  • Observation

Reasoning: based on scientific principles

  • Each piece of evidence may have a different justification for why it supports the claim

Descent With Modification:

Evolution change is based on the interactions between populations and their environments which results in adaptations (inherited characteristics) to increase fitness

Evolution:

  1. Descent with modifications (Darwin)

  2. Change overtime in the genetic composition of a population from generation to generation

Aristotle:

  • Species are fixed (unchanged)

  • Scala naturae - Life forms arrange on ladder of increasing complexity

Old Testament (Creationism):

  • Earth - 6000 years old

  • Perfect species individually designed by God for a particular purpose

Carolus Linnaeus - Founder of Taxonomy:

  • Binomial nomenclature

    • Domain - Kingdom - Phylum - Class - Order - Family - Genus - Species

    • Dear - King - Philip - Came - Over - For - Good - Spaghetti

Humans:

D - Eukarya

K - Animalia

P - Chordata

C - Mammalia

O - Primata

F - Hominidae

G - Homo

S - Homo Sapien

Cuvier:

  • Paleontologist - Studies fossils

  • Deep strata (rock layers) - Very different fossils from current life

  • Opposed idea of evolution

  • Boundaries between strata = Many living species destroyed by catastrophic event, then repopulated by immigrant species

  • Hutton - Geologic change results from slow and gradual, continuous processes

  • Lyell - Earth’s processes same rate in past and present

    • Earth is very old

  • Slow and subtle changes in organisms - Big Change

Lamarck:

  • Published theory of evolution (1809)

  • Use and disuse - Parts of body used bigger, stronger

  • Inheritance of acquired characteristics - modifications that can be passed on

  • Importance - Recognized that species evolve, although explanation was flawed

Malthus:

  • More babies born than deaths

  • Consequences of overproducing within environment = war, famine

  • Struggle for existence

Darwin:

  • English naturalist

  • 1831 - Joined the HMS Beagle for a 5-year research voyage around the world

  • Collected and studied plant and animal specimens, bones, fossils

  • Notable Stop - Galapagos Islands

  • Waited 30 years before publishing his ideas on evolution

  • Alfred Russell Wallace - Published paper on natural selection first (1859)

  • Mechanism for evolution is natural selection

  • Didn’t use ‘evolution’ but rather ‘descent with modification’

Natural Selection:

Adaptations enhance an organism’s ability to survive and reproduce in specific environments

  • Desert Fox - Large Ears, Arctic Fox - Small Ear

If humans can create substantial change over short time, nature can stay over long time

Homology:

  • Characteristics in related species that can have underlying similarity even though functions may differ

    • Homologous Structures - Similar anatomy from common ancestors

    • Embryotic Homologies - Similar early development

    • Vestigial Organs - Structures with little or no use

    • Molecular Homologies - Similar DNA and amino acid sequences

Convergent Evolution:

  • Distantly related species can resemble one another

    • Similar problems similar solutions

  • Analogous Structures: Similar structures, function in similar environments

    • Eg: Torpedo shape of shark, penguin, dolphin

Fossil Record:

  • Fossils - Remains or traces of organisms from past

  • Found in sedimentary rock

  • Paleontology - Study of fossils

  • Show evolutionary changes that occur over time and origin of major new groups of organisms

Pinky - Shrink

Ring - Nonrandom mating

Middle - Mutation

Index - Movement

Thumb - Adaptations

Hardy-Weinberg Equilibrium:

  1. Large Population

  2. Random Mating

  3. No Mutations

  4. No Movement

  5. No Adaptations

  • R - Dominant Allele

  • r - Recessive Allele

  • RR - Homozygous Dominant

  • Rr - Heterozygous

  • rr - Homozygous Recessive

P + Q = 1        P = R         P² = RR        Q = r        Q² = rr

P² + 2PQ + Q² = 1         2PQ = Rr

  • In nature it is not likely all conditions for H-W Equilibrium will be met - Populations are evolving

  • Allele/genotype frequency changes due to mutations and nonrandom mating are minor

  • Three major mechanisms of evolution

    • Natural Selection

    • Genetic Drift

    • Gene Flow

Mechanisms of Microevolution:

  1. Natural Selection - Differential reproductive success

  2. Genetic Drift - Unpredictable fluctuation of alleles from one generation to next

  • Significant genetic drift in small populations

  • Allele frequencies change at random

  • Can lose genetic variation in populations

  • Can cause harmful alleles to become fixes

  • Types

    • A. Founder Effect - Certain alleles over/underrepresented

    • B. Bottleneck Effect - Severe drop in population size

Types of Selection:

Directional - 1 of the extremes is favored

Stabilizing - Extremes are selected against - Middle is favored

Disruptive - Extremes are favored - Middle is selected against

Balancing Selection:

Diploidy:

  • Inherit 2 alleles

  • Recessive alleles hidden in heterozygotes

Heterozygote Advantage:

  • Heterozygotes have better survival

    • Eg: Heterozygotes for Sickle Cell Anemia protected against Malaria

Sexual Selection:

  • Certain individuals are more likely to obtain mates

  • Sexual Dimorphism - Difference between 2 sexes

  • Intrasexual Selection - Competition within the same sex

  • Intersexual Selection - Mate choice

Type of Evolution:

Gradualism:

  • Natural selection gradually changes the average features of a species

  • This process continues for long enough time for a species to change into a new species and the original species becomes extinct

Punctuated Equilibrium:

  • Periods of rapid speciation followed by long periods of stasis (no change)

Divergent:

  • One species evolved into 2 different species

  • Results in homologous structures

Convergent:

  • 2 separate species in different areas evolve to look or behave in similar manner

  • Results in analogous structures

Coevolution:

  • 2 species that have a partnership or symbiotic relationship

  • Evolve together to maintain the relationship

Natural Selection cannot make perfect organisms

1) Selection can only edit existing variations

2) Evolution is limited by historical constraints

3) Adaptions are often compromises

4) Chance, natural selection, and the environment interact

Phylogeny/Tree of Life:

  • Phylogeny - Evolutionary history of a species or group of related species usually organized into a phylogenic tree

Biological Species Concept:

  • Species - Population or group of populations whose members have the potential to interbreed in nature and produce viable, fertile offspring

    • Reproductively compatible

  • Reproductive Isolation - Barriers that prevent members of 2 species from producing viable, fertile hybrid

Prezygotic: Prevent mating or hinder fertilization

  • Habitat Isolation

  • Temporal Isolation

  • Behavioral Isolation

  • Mechanical Isolation

  • Gametic Isolation

Postzygotic: Prevent hybrid zygote from developing into fertile adults

  • Reduced hybrid viability - Miscarriage

  • Reduced hybrid fertility - No more mules

  • Hybrid breakdown - Smaller/weaker

Species

  • Morphological - By body shape, size, and other structural features

  • Ecological - Niche/role in community

  • Phylogenic - Share a common ancestor, form one branch on tree of life

  • Allopatric - Geographically Isolated

    • Caused by geological events 

    • Evolves by natural selection    

      • Ex: Squirrels on N/S rims

  • Sympatric - Overlapping populations within same geographic area 

    • Gene flow between subpopulations blocked by:

      • Polyploidy

      • Habitat differentiation

      • Sexual selection

        • Ex: Polyploidy in 80% of plants

Cells:

Prokaryote - Domain bacteria and archaea

  • Before

  • No nucleus

  • NDA in a nucleoid

  • Cytosol

  • No organelles other that ribosomes

  • Small size

  • Primitive

  • Ex: Bacteria

Eukaryote - Protists, Fungi, Plants, Animals

  • True

  • Has nucleus and nuclear envelope

  • Cytosol

  • Membrane-bound organelles with specialized structure and function

  • Much larger in size

  • More complex

  • Ex: Plant and animal cell

  • Cells must be small to maintain a large surface area to volume ration

  • Large surface area allows higher rate of chemical exchange between cell and environment

Membrane Structure and Function:

Cell membrane - Selectively permeable

  • Somethings in and out

  • Fluid mosaic model

    • Things can move

    • Made up of multiple components

      • Phospholipid

      • Proteins

      • Carbohydrates

      • Cholesterol

Diffusion - Movement from a high concentration to a low concentration

Simple Diffusion: Type of passive transport

  • No energy required

Equilibrium: State of balance (homeostasis)

Facilitated Diffusion: Channel/carrier protein

Active Transport:

  • Uses energy (ATP)

  • Low to high

  • Creates concentration gradient

Early Membrane Model:

  • (1935) Davison/Danielli - Sandwich model

  • Phospholipid bilayer between 2 protein layers

  • Problems - Varying chemical composition of membrane, hydrophobic protein parts

Membrane:

  • Low temps - Phospholipids with unsaturated tails (Kinks prevent close packing)

  • Cholesterol resists changes by:

    • Limit fluidity at high temps

    • Hinder close packing at low temps

  • Adaptations - Bacteria in hot springs (unusual lipids); winter wheat (high unsaturated phospholipids)

Membrane Proteins:

Integral Proteins:

  • Embedded in membrane

  • Determined by freeze fracture

  • Transmembrane with hydrophilic heads/tails and hydrophobic middles

Peripheral Proteins:

  • Extracellular or cytoplasmic sides of membrane

  • Not embedded

  • Held in place by the cytoskeleton or ECM

  • Provides stronger framework

Carbohydrates:

  • Function - Cell to cell recognition; developing organisms

  • Glycolipids, glycoproteins

  • Ex: Blood transfusions

Selective Permeability:

  • Small nonpolar molecules cross easily: hydrocarbons, hydrophobic molecules, CO2, O2, N2

  • Polar uncharged molecules. including H2O

    • Pass in small amounts

  • Hydrophobic core prevents passage of ions, large polar molecules - movement through embedded channel and transport proteins

Passive Transport:

  • No energy (ATP) needed

  • Diffusion down concentration gradient (high to low)

  • Ex: Hydrocarbons; CO2, O2, H2O

Water Potential:

H2O moves from high potential to low potential

  • Water potential - Free energy water

  • Solute potential - concentration (osmotic potential)

  • Pressure potential - Physical pressure on solution; turgor pressure

    • Pure water - MPa

    • Plant cells - 1MPa

Calculate Solute Potential:

  • -iCRT

  • i - Ionization constant (# of particles made in water)

  • C - Molar concentration

  • R - Pressure constant (0.0831 liter bars/mole-k)

  • T - Temperature in K (273 + C)

  • The addition of solute to water lowers the solute potential (more negative) and therefore decreases the water potential

Water Movement:

From and area of:

  • Higher potential to lower potential

  • Low solute concentration to high solute concentration

  • High pressure to low pressure

Facilitated Diffusion:

Transport Proteins - (Cannel or carrier proteins) help hydrophilic substances cross

  • Two ways:

    • Provide hydrophilic channel

    • Loosely bind/carry molecule across

  • Ex - Ions, polar molecules (H2O Glucose)

Aquaporin - Channel protein that allows passage of H2O

Active Transport:

  • Requires energy (ATP)

  • Proteins transport substances against concentration gradient (low to high)

    • Ex - Na+/K+ pump, proton pump

Electrogenic Pump - Generate voltage across membrane

Cotransport - Membrane protein enables “downhill” diffusion of one solute to drive “uphill” transport of other

  • Ex - Sucrose - H+ cotransporter (sugar loading in plants)

Passive:

  • Little or no energy

  • High to low

  • Down the gradient

  • Ex - Diffusion, Osmosis, Facilitated Diffusion

Active:

  • Requires energy

  • Low to high

  • Against the gradient

  • Ex - Pumps, exo/endocytosis

Osmoregulation:

  • Control solute and water balance

  • Contractile vacuole - “Bilge Pump” forces out fresh water as it enters by osmosis

  • Ex - Paramecium Caudatum - Freshwater protist

Bulk Transport:

  • Transport of proteins, polysaccharides, large molecules

Endocytosis - Take in macromolecules and particulate matter, form new vesicles from plasma membrane

Exocytosis - Vesicles fuse with plasma membrane, secrete contents out of cell

:

  • The highly complex organization of living systems requires constant input of energy and the exchange of macromolecules

  • Organisms use energy to maintain organization, reproduce, and grow

Metabolism:

The totality of an organism’s chemical reactions

  • Manage the materials and energy resources of a cell

  • Metabolic Rate - Total amount of energy an animal uses in a unit of time

  • In general, the smaller the organism, the higher the metabolic rate

  • Catabolic Pathways - Release energy by breaking down complex molecules into simpler compounds

    • Ex - Digestive enzymes break down food and release ATP

  • Anabolic Pathways - Consume energy to build complex molecules from simpler ones

    • Ex - Amino acids link to form muscle protein

  • Kinetic Energy - Associated with motion

    • Heat is KE associated with random movement of atoms or molecules

  • Potential Energy - Stored energy as a result of its position or structure

    • Chemical Energy is PE available for release

  • Energy can be converted from one form to another

    • Ex - Chemical to mechanical to electrical

Thermodynamics:

  • Study of energy transformations that occur in matter

  • Closed system - Isolated from its surrounding

  • Open system - Energy and matter can be transferred between the system and its surroundings

  • Organisms - Open systems

    • A net gain in energy results in energy storage or the growth of an organism

    • A new loss of energy results in loss of mass and/or death of an organism

  • A living cell isn’t at equilibrium

    • Constant flow of materials

  • Three kinds of work

    • Mechanical

    • Transport

    • Chemical

  • Cells manage energy resources to do work by energy coupling - using an exergonic process to drive an endergonic one

  • ATP (Adenosine Triphosphate) is the cell’s main energy source in energy coupling

  • ATP - Adenine Ribose 3 phosphates

  • Hydrolysis - Energy released

ATP Performance:

  • Exergonic release of Pi is used to do the endergonic work of cell

  • When ATP is hydrolyzed it becomes ADP

  • Catalyst - Substance that can change the rate of reaction without being altered in the process

  • Enzyme - Biological catalyst

  • Speeds up metabolic reactions by lowering the activation energy

Substrate Specificity:

  • The reactant that an enzyme acts on is called the enzyme’s substrate

  • The enzyme binds to its substrate forming an enzyme-substrate complex

  • The active site is the region on the enzyme where the substrate binds

  • Induced Fit - Enzyme fits snugly around substrate - “Clasping handshake”

  • An enzymes activity can be affected by temperature, pH, or chemicals

Structure/Function:

  • Change to the molecular structure of a component in an enzymatic system may result in a change of function or efficiency of the system

  • Denaturation - Disrupt protein structure

    • Reduce enzymatic activity

  • Environmental pH - Alter efficiency of enzyme activity - disrupts of H-Bonds

Cofactors: Non-protein enzyme helpers such as minerals (Zn, Fe, Cu)

Coenzymes: Organic cofactors (Vitamins)

  • Enzyme Inhibitors -

    • Competitive inhibitors - Binds to the active site of an enzyme, competes with substrate

    • Noncompetitive Inhibitors - Binds to another part of an enzyme → enzyme changes shape → active site is nonfunctional

Regulation:

  • To regulate metabolic pathways, the cell switches on/off the genes that encode specific enzymes

  • Allosteric Regulations - Proteins function at one site is affected by binding of a regulatory molecule to a separate site

  • Activator - Stabilizes active site

  • Inhibitor - Stabilizes inactive form

  • Cooperativity - One substrate triggers shape changes in another active site

Cellular Respiration:

  • Glycolysis

  • Krebs/Citric Acid Cycle

  • Electron Transport Chain

  • E flows into ecosystem as sunlight

  • Autotrophs transform it into chemical E (O2 replaced)

  • Cells use some of chemical E in organic molecules to make ATP

  • E leaves as heat

Energy Harvest:

  • Energy is released as electrons “fall” from organic molecules to O2

  • Broken down into steps -

  • Food (Glucose) NADH+ETC O2

  • Coenzyme NAD+ = Electron acceptor

  • NAD+ picks up 2e and 2H+ NADH (Stores E)

  • NADH carries electrons to the electron transport chain

  • ETC - Transfers e to O2 to make H2O - releases energy

Glycolysis:

  • “Sugar splitting”

  • Believed to be ancient (early prokaryotes - no O2)

  • Occurs in cytosol

  • Partially oxidizes glucose (6C) to 2 pyruvates (3C)

  • Net gain - 2ATP + 2NADH

    • 2 H2O

    • No O2 required

Stage 1: Energy Investment

  • Cells use ATP to phosphorylate compounds of glucose

Stage 2: Energy Payoff

  • Two 3C compounds oxidized

  • For each glucose molecule

    • 2 net ATP

    • 2 molecules of NAH+ NADH

Substrate - Level Phosphorylation:

  • Generate small amount of ATP

  • Phosphorylation - Enzyme transfers a phosphate to other compounds

Pyruvate Oxidation:

  • Pyruvate Acetyl CoA

  • CO2 and NADH produced

Krebs Cycle:

  • Occurs in mitochondrial matrix

  • Acetyl CoA, Citrate, and CO2 released

  • Net gain - 2 ATP, 6NADH, 2FADH2 (electron carrier)

  • ATP produced by substrate - level phosphorylation

Electron Transport Chain (ETC):

  • Collection of molecules embedded in inner membrane of mitochondria

  • Tightly bound proteins + non-protein components

  • Alternate between reduced / oxidized states as accepts / donate e

  • Does not make ATP directly

  • Ease falls of e from food to O2

  • 2H+ + ½ O2 → H2O

Photosynthesis:

  • Plants and other autotrophs are producers of biosphere

  • Photoautotrophs - Use light E to make organic molecules

  • Heterotrophs - Consume organic molecules from other organisms for E and carbon

  • Photosynthesis - Converts light energy to chemical energy of food

  • Chloroplasts - Sites of photosynthesis in plants

  • Stomata - Pores in leaf (CO2 enters / O2 exits)

Sites of Photosynthesis:

  • Mesophyll - Chloroplasts mainly found in these cells of leaf

  • Chlorophyll - Green pigment in thylakoid membranes of chloroplasts

  • 6CO2 + 6H2O + Light Energy → C6H12O6

  • Redox Reaction - Water is split → E- transferred with H+ to CO2 → sugar

    • Oxidation → lose E-

    • Reduction → gain E-

  • Photosynthesis = Light reaction + Calvin Cycle

Nature of Sunlight:

  • Light = Energy = Electromagnetic radiation

  • Shorter Wavelength - Higher E

  • Visible Light - Detected by human eye

  • Light - Reflected, transmitted, or absorbed

Photosynthetic Pigments:

  • Pigments absorb different wavelengths of light

  • Chlorophyll - Absorbs violet blue / red light, reflect green

    • Chlorophyll A (Blue-Green) - Light reaction, converts solar to chemical E

    • Chlorophyll B (Yellow-Green) - Photoprotection, broaden color spectrum

  • Absorption Spectrum - Determines effectiveness of different wavelengths for photosynthesis

Calvin Cycle:

  • Uses ATP and NADPH to convert CO2 to sugar

  • Produces 3-C sugar G3P

  • Three phases

    • Carbon fixation

    • Reduction

    • Regeneration of RuBP (CO2 acceptor)

Photorespiration:

  • Metabolic pathway which -

    • Uses O2 and produces CO2

    • Uses ATP

    • No sugar production (Rubisco binds O2 → breakdown of RuBP)

  • Occurs in hot, dry bright days which stomata close (Conserve H2O)

  • Early atmosphere - Low O2, high CO2

Evolutionary Adaptations:

  • Problem with C3 plants

    • Co2 fixed to 3-C compound in Calvin Cycle

      • (Ex: Rice, Wheat)

    • Hot, dry days

      • Partially close stomata, lower Co2

      • Photorespiration

      • Lower photosynthetic output (No sugar made)

  • C4 plants

    • Co2 fixed to 4-C compound

      • (Ex: Corn, Sugarcane, grass)

    • Hot, dry days → Stomata close

      • 2 cell types - Mesophyll / Bundle sheath cells

      • Mesophyll - PEP carboxylase fixes CO2

      • Bundle Sheath - CO2 used in Calvin Cycle

  • CAM Plants

    • Night - Stomata open → CO2 enters → converts to organic acid, stored in mesophyll cells

    • Day - Stomata closed → Light reactions supply ATP, NADPH; CO2 released from organic acids for Calvin Cycle

      • (Ex: Cacti, Succulent, Pineapple)

3 Phases of Signal Transduction:

  • Reception - The target cell’s detection of a signal molecule coming from the outside

  • Transduction - The conversion of a signal to a form that can bring a specific cellular response

  • Response - The specific cellular response to the signal molecule

Ligand:

  • Signaling molecule

    • Ex: Proteins, amino acid, steroids

Receptor:

  • Protein that detects specific ligands

  • Lock and key type fit

Types of Cell Signaling:

  • The type of signaling a cell uses is based on the distance between the cell it is trying to signal

4 main types:

  • Juxtracrine / Direct Communication

    • Cells are touching. One cell can recognize the molecules on the adjacent cell

      • Plant Ex: Plasmodesmata

      • Animal Ex: Gap Junctions

      • Orchestrates early embryo development

  • Paracrine

    • Ligands produced by cells can travel through extracellular fluid (diffusion) and can be read by other local cells

      • Short-lived molecules

      • Two outcomes:

        • Read by another cell

        • Degraded by enzymes

  • Endocrine

    • Ligand released by a cell and makes its way to the circulatory system

      • Can be spread to the enire body

      • Long-lived molecules known as hormones

      • Used extensively in plants and animals

  • Synaptic

    • Rapid communication with distant cells using nerve cells’ long fiber-like extensions

      • Ligands are called neurotransmitters

      • Chemical synapse - Association of the neuron and its target cells

      • Used by the nervous system

        • Ex: Touch

  • Autocrine signaling is another important signaling event

    • Occurs when a cell’s signals itself. Production and secretion of an extraellular mediator by a cell followed by binding of that mediator by receptor on the same cell to initiate signal transduction

      • Ex: Cancer cells making their own growth hormone rather than relying on its release from the pituitary gland

Cell Junctions:

  • Tight junctions - Belts around the epithelial cells that line organs and serve as a barrier to prevent leakage into or out of those organs

  • Plasmodesmata - Connect one plant cell to the next. They are analogous to gap junctions in animal cells

Feedback Regulation:

  • Negative Feedback Loop - Inhibits a response by reducing the initial stimulus, thus preventing excessive pathway activity

  • Positive Feedback Loop: Reinforces a stimulus to produce an even greater response

  • Desmosomes - ‘Spot Welds’ found in many tissues that are subjected to severe mechanical stress such as skin epithelium or the neck of the uterus, which must expand greatly during childbirth

  • Gap Junctions - Permit the passage of materials directly from the cytoplasm from one cell to the cytoplasm of an adjacent cell. In the muscle tissue of the heart, the flow of ions through the gap junctions coordinates the contractions of the cardiac cells.

  1. Reception:

  • Occurs when a signal molecule (Ligand) binds to a receptor protein

  • Found in 2 places:

    • Plasma membrane

      • Binds to water-soluble ligands

    • Intracellular

      • Found inside plasma membrane in cytoplasm or nucleus. The ligand crosses the membrane (it’s hydrophobic), like the steroid hormone testosterone

Cell Cycle: Life of a cell from its formation until it divides into 2 cells

Functions of Cell Division:

  • Reproduction

  • Growth and development

  • Tissue renewal

Genome - All of a cell’s genetic info (DNA)

  • Prokaryote - Single, circular chromosome

  • Eukaryote - More than one linear chromosome

    • Ex: Human - 46, Mouse - 40, Fruit fly - 8

  • Each chromosome must be duplicated before cell division

Duplicated Chromosome - 2 sister chromatids attached by a centromere

Somatic Cells:

  • Mody cells

  • Diploid (2n) - 2 of each type of chromosome

  • Divide by mitosis

  • Humans - 2n = 46

Gametes:

  • Sex cells (Sperm / Egg)

  • Haploid (n) - 1 of each type of chromosome

  • Divide by mitosis

  • Humans - n = 23

Checkpoint: Control point where stop / go signals regulate the cell cycle

Major Checkpoints:

  1. G1 (Most important)

  • Controlled by cell size, growth factors, environment

  • ‘Go’ → Completes whole cell cycle

  • ‘Stop’ → Cell enters nondividing state (Go phase)

    • Nerve muscle cells stay at go; liver ells called back from go

  1. G2

  • Controlled by DNA replication completion, DNA mutations, and cell size

  1. M-Spindle

  • Check spindle fiber (microtubule) attachment to chromosomes at kinetochores (anchor sites)

Kinetochore - Protein associated with DNA at centromere

Kinases - (Cyclin-dependent kinase, CDK) Protein enzyme controls cell cycle; active when connected to cyclin

Cyclin - Proteins which attach to kinases to activate them; levels fluctuate in the cell cycle

External Regulator Factors:

  • Growth - Proteins released by other cells to stimulate cell division

  • Density Dependent Inhibition - Crowded cells normally stop dividing; cell-surface protein binds to adjoining cell to inhibit growth

  • Anchorage Dependence - Cells must be attached to another cell or ECM (extracellular matrix) to divide

Transformation:

  • Process that converts a normal cell to a cancer cell

    • Tumors - Mass of abnormal cells

    • Benign - Lump of cells remain at original site

    • Malignant - Invasive and impairs functions of 1+ organs

    • Metastasis - Cells separate from tumor and travel to other parts of the body

G Protein-Coupled Receptor:

  • A membrane receptor that works with the help of G protein

  1. The ligan or signaling molecule has bound to the G protein-coupled receptor. This causes conformational change in the receptor so that it may now bind to an inactive G protein, causing a GTP to displace the GDP. This activates the G protein.

  2. G protein binds to a specific enzyme and activates it. When the enzyme is activated, it can trigger the next step in a pathway leading to a cellular response. All the molecular shape changes are temporary. To continue the cellular response, new signal molecules are required.

Intracellular Receptors:

  • Bind hydrophobic ligands

    • Hydrophobic ligands can easily cross the plasma membrane

  • Main class of intracellular receptors are nuclear receptors

Insulin / Glucagon:

  • Insulin (decreases blood sugar) and glucagon (increases blood sugar) are antagonistic hormones that help maintain glucose homeostasis

  • Pancreas has clusters of endocrine cells called pancreatic islets with alpha cells that produce glucagon and beta cells that produce insulin

Target Tissue:

  • Insulin reduces glucose levels by

    • Promoting cellular uptake of glucose

    • Slowing glycogen breakdown in the liver

    • Promoting fat storage, not breakdown

  • Glucagon increases glucose levels by

    • Stimulating conversion of glycogen to glucose in the liver

    • Stimulating the breakdown of fat and protein into glucose

Phases of Cell Cycle:

  • The mitotic phase alternates with interphase

    G1 → S → G2 → Mitosis → Cytokinesis

  • Interphase (90% of cell cycle)

    • G1 - Cell grows and carries out normal functions

    • S - Duplicates chromosomes (DNA replication)

    • G2 - Prepares for cell division

  • M Phase (Mitotic)

    • Mitosis - Nucleus divides

    • Cytokinesis - Cytoplasm divides

Mitosis:

    Prophase → Metaphase → Anaphase → Telophase

  • Continuous process with observable structural features

    • Chromosomes become visible (Prophase)

    • Alignment at the equator (Metaphase)

    • Separation of sister chromatids (Anaphase)

    • Form 2 daughter cells (Telophase and Cytokinesis)

Cytokinesis:

  • Cytoplasm of cell divided

  • Animal Cells - Cleavage furrow

  • Plant Cells - Cell plate forms