Biology II: Exam 3

Organization of Animal Bodies

  • all animal cells share similarities in which they…

    • lack cell walls (instead have plasma membrane; gatekeepers)

    • exchange materials with surroundings

    • obtain energy from organic nutrients

    • synthesize complex molecules

    • reproduce themselves

    • detect and respond to signals in their immediate environment

  • levels of organization

    • Cellular - Phylum Porifera

    • Tissue - Phylum Cnidaria and Phylum Ctenophora

    • Organ - Phylum Platyhelminthes

    • Organ Systems - Phylum Annelida, Phylum Arthropoda, Phylum Mollusca, Phylum Echinodermata, Phylum Chordata

  • internal organization

    • cells with similar properties group together to form tissues

    • tissues combine together to form organs

    • organs are linked together to form organ systems

Tissues

  • tissues are an association of many cells that have similar structure

    • epithelial tissue

    • connective tissue

    • nervous tissue

    • muscle tissue

  • Epithelial Tissue

    • sheets of densely packed cells that…

      • covers body and encloses organs

      • lines walls of body cavities and organs

    • specialized to protect and secrete/absorb ions and organic molecules

    • cells have a variety of shapes

      • based on shape of cell

        • cuboidal - square shaped cells

        • squamous - flat-like cells

        • columnar - rectangular shaped cells

      • based on layers of cells

        • simple - one layer of cells

        • stratified - multiple layers of cells

        • pseudostratified - one layer, but looks like multiple

    • all epithelial cells are asymmetrical or polarized

      • one side of the cell rests on the basal lamina (basement membrane) and the other faces the environment

    • types of epithelial tissues

      • simple squamous - single layer of flat cells

      • simple cuboidal - single layer of square cells

      • simple columnar - single layer of rectangular cells

      • pseudostratified columnar - one cell thick with all cells connected to base membrane

      • stratified squamous - multiple layers of flat cells

      • transitional - stretchable tissue in urinary bladder

  • Connective Tissue

    • connects, surrounds, anchors, binds, transports, and supports

    • form extracellular matrix (ECM) around cells

      • provides scaffold for attachment

      • protects and cushions

      • mechanical strength

      • transmits information

    • types of connective tissue

      • blood - transports and protects

      • adipose (fat) - insulation, energy, support, and protection

      • bone - support and protection

      • cartilage - support and flexibility

      • loose - holds internal organs in place

      • dense - strength and support

  • Muscle Tissue

    • specialized cells that generate mechanical force

    • skeletal muscle

      • attach to bone or exoskeleton for locomotion

      • elongate fibers

      • voluntary control

    • smooth muscle

      • surrounds tubes and cavities for propulsion of contents

      • flattened cells

      • involuntary control

    • cardiac muscle

      • only in the heart

      • elongates fibers

      • involuntary control

  • Nervous Tissue

    • a complex network of neurons (nerve cells) and other cells

    • initiate and conduct electrical signals from one part of the animal body to another

    • electrical signals produced in one neuron may stimulate or inhibit other neurons to…

      • initiate new electrical signals

      • stimulate muscle cells to contract

      • stimulate glandular cells to release chemicals

    • contains neuroglial cells (more numerous than neurons)

      • provide metabolic support, maintenance, ion balance, and cleaning for the neurons

      • produces new glial cells and neurons

        • neurons CAN’T produce new neurons

Organs

  • composed of two or more of the major types of tissues

    • may form sheets, tubes, layers, bundles, or strips

  • organ systems - different organs that work together to perform an overall function

    • frequently work with other organ systems

    • EX. nervous, circulatory, and endocrine systems functions to influence how much water kidneys retain

  • spatial arrangement of organs into organ systems part of overall body plan

    • body plan controlled by highly conserved family of genes with homologs in all animals (Hox Genes) especially in vertebrate

Organisms

  • structure and function

    • organization of structure can usually predict the functions

      • EX. big structure, more complex functions

    • concentrate on increasing complexity of structure, thus increasing complexity of organismal functioning

    • mostly emphasis on vertebrate

  • Homeostasis

    • changing variables in environment

      • air temperature

      • water temperature

      • food supply

      • water supply

      • pH

      • O2 concentration

    • homeostasis is the process of adjusting to external environment and maintaining a stable internal environment

      • there is no set form of home; all animals may have a different versions

Integumentary System (Vertebrate Integuments)

  • skin and all other accessories

    • skin is the largest organ in vertebrates

      • consists of two layers

        • epidermis (outer)

        • dermis (inner)

  • Functions

    • protection from absorption

    • protects against water loss

    • barrier to disease

    • (little) protection from UV light

    • temperature regulation

    • contains sensory receptors (EX. touch pain)

    • (limited) excretion

  • Epidermis

    • thinner, outer layer

    • nutrients diffuse into here from the dermis

    • stratified squamous epithelium

    • most become dead cells from not enough nutrients

    • cell types

      • Langerhans Cells

        • defensive cells

      • Melanocytes

        • produces melanin pigment

        • skin coloration

        • protect from UV

      • Merkel Cells

        • touch receptors

      • Keratinocytes

        • PRIMARY cell type

        • produces insoluble protein keratin

        • amount of keratin increase from inside to out (more keratin on outside)

        • keratin fills cytoplasm and impairs nutrient diffusion causing cells to die

  • Dermis

    • inner layer of skin

    • thicker than epidermis

    • highly vascularized (blood vessels)

    • contains sensory structures, vessels, nerves, glands

    • origin of hair, feathers, scales

    • sensory structures

      • Meissner’s Corpuscles

        • light touch

      • Pacinian Corpuscles

        • deep pressure

        • vibration

  • Sweat Glands

    • temperature regulation

    • produces sweat (primarily water)

    • 2.5 million glands in body

    • releases heat

  • Sebaceous Glands

    • all over body EXCEPT palms and soles

    • large on face, neck, and upper chest

    • produces sebum (oil); lubricates and softens hair and skin

      • aquatic animals have more sebum

  • Hypodermis (Subcutaneous Layer)

    • lies beneath dermis

    • NOT a layer of skin

    • contains adipose (fat) tissue

    • functions

      • body contour

      • insulation (holds in heat)

      • energy storage

    • females have thicker layers of adipose than males (softer skin)

Digestive System

  • Intro to nutrition

    • nutrients - any substances taken in by organisms that is needed for survival, growth, develop, maintenance, or reproduction

    • nutrition - process of consuming and using food and nutrients

    • animals get nutrients from consuming food

  • animals are heterotrophs

    • CAN’T manufacture own food

    • require synthesized organic compounds of plants or other animals to supply materials for…

      • survival

      • growth

      • maintenance

      • reproduction (highest amount of food/energy intake)

  • Gut Tracts

    • Blind Gut

      • NO CAVITY between gut and body wall

      • one opening

      • primitive form (jellyfish, flatworms)

    • Tube-within-a-Tube

      • flow through digestive tube

      • body cavity between gut and body wall

      • separate openings

      • allows for expansion

  • Digestive Enzymes (require H2O)

    • carbohydrates

    • proteases (proteins)

    • lipases (lipids)

    • nucleases (nucleic acids)

  • food processing in animals

    • occurs in five phases

      1. Ingestion - food taken into body and moves into digestive cavity; usually called alimentary canal

      2. Digestion - food is broken down into smaller molecules

      3. Transportation

      4. Absorption - ion, water, and small molecules diffuse or are transported into circulatory system

      5. Egestion - undigested materials and other wastes are passed from the body

  • alimentary canal

    • digestive tract or tube (GI Tract)

    • single, elongated tube with opening at both ends

    • contains smooth muscle in cell walls

    • lined by simple epithelial tissue

      • synthesizes and secrete digestive enzymes

      • secrete hormones

      • transport digested materials

    • several specialized regions

      • different environments for different processes

      • storage area

    • structure of GI Tract

      • same general structure from midpoint of esophagus to anus (cloaca)

        • lumen lined by epithelium and gland cells

        • secretory cells release protective layers of mucus

        • other cells release hormones

        • glands release acid, enzymes, water, and ions

      • epithelial cells lined by tight junctions and surrounded by layers of tissue made of smooth muscles, neurons, connective tissue, and blood vessels

        • neurons activated by sight and smell of food, presence of food in tract

  • 5 Regions of Alimentary Canal

    • Region of Reception

      • buccal cavity (mouth and accessory organs)

        • ingestion site and digestion site (chemical and mechanical)

        • jaws, teeth, muscles, tongues, salivary glands (saliva —> enzymes)

        • pharynx (back of mouth cavity)

          • where digestive and respiratory cross paths

    • Region of Conduction

      • esophagus

        • tube carrying materials from mouth to rest of alimentary canal

        • from neck region to the chest

        • conducts food from pharynx to stomach

        • peristalsis - rhythmic, wave-like contraction which propels food forward in GI Tract

          • squeezes food along the tract like toothpaste

    • Region of Storage and More Digestion

      • stomach (mostly)

        • muscular sac-like organ involved in storing food

        • muscle nature helps break up food; partial protein digestion

        • regulates rate of emptying into small intestine

      • secretions

        • hydrochloric acid - kills microbes; dissolves particulate matter (Parietal cells)

        • pepsinogen - converted to pepsin to begin protein digestion (Chief cells)

      • epithelium coated with alkaline mucus

      • lumen (cavity) stomach

      • pepsinogen + HCl —> pepsin (protein breakdown)

    • Region of Terminal Digestion and Absorption of Food

      • small intestine

      • nearly all digested food and absorption of food

      • hydrolytic enzymes found on apical surface of epithelial cells or secreted by pancreas into lumen

      • products of digestion absorbed across epithelial cells and enter blood

        • vitamins/minerals and water absorbed (glucose)

      • small intestine length = 8x height of organism

      • small intestine specialized to carry out bulk of digestion and absorption

        • mucosal infoldings

        • villi - finger-like projections

          • epithelial cells with microvilli create brush border (extensions of plasma membrane)

      • specializations increases surface areas 600-fold

        • increase likelihood of food particles encountering digestive enzymes and being absorbed

      • 3 sections of small intestine

        • duodenum

        • jejunum

        • ileum

      • surface modifications that increase surface area for absorption

        • plicae circulares

          • large folds of epithelial lining

          • increase surface area 2-3 times

        • villus (villi)

          • finger-like projections

          • increases surface area 10 times

          • lined with epithelium

        • microvilli

          • folding of plasma membrane of cells lining the villus

          • increase 20 times

    • Region of H2O Absorption

      • large intestine

        • mammals - about 1.5 meters in length

        • lack plicae, villi, microvilli

        • H2O absorbed through epithelium

        • function - humans (~1400 mL of H2O/day)

        • compacts and solidates all eliminated feces

      • waste

        • water - 75%

        • inorganic substances - 5%

        • roughage - 8%

        • fat - 5% (very hard to break down)

        • undigested protein, dead cells, bile - 2%

      • most mammals end of large intestine is the anus

      • anus-opening at posterior end of alimentary canal for release of waste production

      • cloaca - some vertebrates; chamber receiving contents of digestive, reproductive, and urinary tracts; vent opening to the outside

  • accessory digestive glands

    • not part of alimentary canal (necessary for proper digestion)

    • salivary glands - secrete saliva

    • pancreas - secrete enzymatic juices through the pancreatic duct into the duodenum

    • liver - produces bile (assists enzymes in break down of large fat globules into smaller forms)

    • bladder - storage of bile; secrete bile into small as needed

  • biomolecules

    • carbohydrates - polysaccharides (starch) —> mono (glucose) & disaccharides (sucrose)

    • proteins - polypeptides —> amino acids (in stomach)

    • fats - glycerol & fatty acids

    • nucleic acids - nucleotides (individual subunits of nucleic acids)

    • *enzyme activity for chemical reaction at biological temperatures (breaks the bonds)

Nervous System

  • Central Nervous System (CNS) - brain and spinal cord in vertebrates

  • Peripheral Nervous System (PNS) - all neurons and their projections outside of CNS

  • *in certain invertebrates with simple nervous systems, the distinction is less clear btwn CNS and PNS

  • cellular components of nervous system

    • nervous system has two classes of cells (neurons and glia)

      • neurons - nerve cells; cells that send and receive electrical and chemical signals to and from each other and other cells throughout body

        • present in ALL animals EXCEPT sponges

        • number of neurons varies as a function of size and behavioral complexity; also varies on brain/body size

      • neuron structure

        • soma (cell body)

          • contains nucleus and organelles

        • dendrites

          • extensions of cell body, single or branching

          • receive incoming signals from nearby neurons

        • axons

          • extension of cell body

          • carries signals to other cells (AWAY from cell body)

          • may have branches and may be wrapped in myelin sheath

          • axon hillock near cell body (base)

          • axon terminals convey electrical or chemical signals to other cells (tips)

          • typically occur in bundles wrapped by connective tissue called nerves

      • glia - surrounds neurons and perform numerous functions (1000x more numerous than neurons)

        • astrocytes

          • metabolic support (nutrition)

          • maintain concentration of ions in extracellular fluid

        • microglia

          • participate in immune functions

          • remove cellular debris

        • myelin sheath

          • interrupted by non insulated nodes of ranvier

          • produced by oligodendrocytes (CNS) and Schwann cells (PNS)

        • radial glial cells

          • warehouse for cellular division

          • can divide into all the other types of glial cells

    • three main types of neurons

      • sensory neurons

        • detect info from outside world or internal body conditions

        • also called afferent neurons —> transmits TOWARDS the CNS

      • motor neurons

        • send signals AWAY from CNS to elicit responses

        • also called efferent neurons

      • interneurons

        • forms interconnections btwn other neurons

        • critical interpretation of info and elicited responses

    • reflex arc

      • stimulus from sensory neurons sent to CNS; little to no interpretation (fewer interneurons)

      • signal transmitted to motor neurons to elicit response (“automatic” signal sent)

        • EX. stimulus —> hand touches stove (receptor)

        • effector —> you move your hand (response)

      • quick and automatic responses

  • electrical properties of neurons

    • membrane potential (“gatekeeper”)

    • ONLY neurons and muscle cells generate electrical signals

    • cells are polarized due to membrane’s permeability

    • difference in ions btwn inside/outside of neuron (causes both chemical and electrical differences); imbalances within glial cells

      • resting membrane potential

        • when neuron is NOT sending signals

        • selectively permeable to cations (+) and anions (-)

        • inside is more (-), outside is more (+)

          • the membrane helps keep the imbalance (creates barrier)

        • anions on inside are drawn to cations on outside, so most ions are near the edge of membrane

  • electrochemical gradient

    • imbalance due to differences inside/outside of neuron

    • chemicals —> K+, Na+, Cl-

    • charge with chemicals + or -

  • 3 factors contributing to resting membrane potential

    1. sodium-potassium pump —> requires ATP

      • pumps 3 Na+ for every 2 K+

    2. ion specific channels allow passive ion movement (high to low)

      • membrane is more permeable (favors) to K

      • K channels more frequently open at resting potential

    3. polarity

      • more (-) inside neuron

  • gated ion channels

    • voltage-gated —> open/close in response to voltage charnes

    • chemical-gated —> open/close in response to chemical changes

  • nerve impulse

    • frequency - language of the message

      • higher frequency, greater excitation level (more immediate response)

      • resting potential —>imbalance btwn K+ and Na+

        • membrane selectively permeable to K, but channels closed for Na and Cl

        • outside neuron 10x more Na, 5x more Cl

        • inside neuron 30x more K

      • action potential - rapid brief change of nerve fibers

        • electric potential of impulse

        • self propagating

        • after passing a given point, membrane returns to resting

        • at given impulse point, channels for Na open and Na diffuses in

        • K is already diffusing out, but increased at impulse

      • sodium-potassium pump

        • complex of proteins in membrane (requires ATP)

        • pumps out Na, carries in K

        • return the imbalance (“normalcy”)

      • nerve impulse rate

        • variable-dependent on complexity of system and organism

          • EX. sea membrane (0.1 m/sec)

          • mammals (120 m/sec) —> nerves move really fast

        • invertebrate animals - speed is related to axon diameter

        • vertebrates - speed is combo of axon diameter and layers of myelin sheath

          • myelin sheath is in sections (Nodes of Ranvier)

          • “saltatorial locomotion” - jumping motion where animal moves a lot (ex. kangaroo)

  • evolution of nervous system

    • Phylum Cnidaria (jellyfish) - simplest neural organization

      • “nerve net” - not a system; simple primitive nerve cells

    • Phylum Platyhelminthes (flatworms)

      • 2 anterior ganglia, each network branching off

      • weak PNS and CNS (no brain or spinal cord)

    • Phylum Annelida (segmented worms)

      • brain, ventral nerve cord, simple motor/sensory neurons

    • Phylum Mollusca (mollusks)

      • squids and octopus may be equal to fish

      • complex - more than earlier groups

    • Phylum Arthropoda (arthropods)

      • similar to annelids and most mollusks

      • EXCEPT “social insects” - well developed brain, complex social behaviors, learning, division of labor, communication (EX. ants)

  • vertebrate nervous system

    • spinal cord and brain = CNS

    • both are surrounded by meninges (layers) - 3 of these

      • dura mater - outer

      • arachnoid - middle

      • pia mater - inner

      • btwn each layer and within canal of spinal cord contains cerebrospinal fluid

    • spinal cord - always dorsal and hollow

      • enclosed within vertebral column

    • brain

      • increase in size with complexity of vertebrate evolution

      • 3 divisions

        • hindbrain - where brain attaches to spinal cord

        • midbrain

        • forebrain - biggest part of the brain

    • brain : spinal cord ratio (ratio increases based on importance of brain)

      • fish —> 2 : 1 (EX. brain is 2x bigger than spinal)

      • amphibians —> 10 : 1

      • reptiles —> 25 : 1

      • birds —> 35 : 1

      • humans —> 55 : 1

      • *birds and mammals have the most complex bodies

Muscular - Skeletal Systems

  • types of animal skeletons

    • a skeleton is a structure that function in support, protection, locomotion (movement from one place to another)

    • three types of skeletons

      • exoskeletons

      • endoskeletons

      • hydroskeletons

        • found in soft-bodied invertebrates that use water pressure for propulsion (EX. jellyfish)

    • exoskeletons

      • internal structures

      • do not protect body surface, only some internal organs

      • found in echinoderms and vertebrates (deuterostomes)

      • minerals provide firmness

      • echinoderm skeletons composed of spiky networks of protein and minerals or mineralized plate-like structures

      • vertebrate skeletons composed entirely of cartilage (cartilaginous fishes) or of cartilage bone

      • advantage - skeleton is internal and grows with the body; composed of living tissue

      • other functions

        • blood cells and platelets are formed in marrow

        • calcium and mineral storage

        • provides attachment sites for skeletal muscle

  • Bone

    • lining, dynamic connective tissue

    • organic components

      • osteoblasts and osteocytes - cells that form bone

      • composed of collagen - triple helical structure provides strength and flexibility

      • osteoblasts - breakdown bone

    • mineral components (salts, phosphates)

    • crystalline mixture of Ca2+, PO4-, and other ions provide rigidity

    • continuously formed, broken down, and reformed

    • formation

      • endochondral - bone replaces cartilage (most bones)

      • intramembranous - bones forms with tissue membranes (skull plates, thin tissue)

    • one cubic inch of bone can sustain 19,000 lbs

    • one mm diameter fiber of collagen can hold 19 lbs of dangling weight

    • types of bone tissue (both provide strength and support)

      • compact bone (ground bone)

        • “dense”

        • composed of osteons

      • spongy bone

        • lacks osteons, lots of air spaces

        • consists of “bony spikes” called trabeculae

    • microscopy anatomy of compact bone

      • osteons - structural unit of spongy bone (doesn’t cover ends of bone)

      • osteonic canal - support and protection from damage, space for arteriole, venules, and nerves

      • trabeculae - spikes of spongy bones

      • periosteum - protective layer around outside of bone

      • lamellae - ring of bone tissues as it develops

      • lacuna - the space between each lamella where osteocytes are located

      • canaliculi - microscopic channels connecting lacunae

    • anatomy of long bone

      • epiphysis - ends of bone (has spongy bone)

      • diaphysis - shaft of bone (mostly compact bone)

      • medullary cavity

        • yellow bone marrow storage (adipose tissue) in diaphysis

        • red bone marrow in epiphysis of certain bones; site of blood cell production

      • compact bone on the outer portion of entire

      • articular surfaces of bone are capped with a layer of hyaline cartilage which acts as a shock absorption

      • periosteum - protective membrane covering external surface of a bone except articular surface

  • the vertebrate system

    • two parts

      • axial skeleton - mid longitudinal axis of body

      • appendicular skeleton - limb bones and glands (everything away from midline)

    • joints

      • formed where two or more bones come together (allow enhanced movement and better contact with substrate)

      • allow movement

        • pivot joints

        • ball-and-socket joints

        • hinge

    • axial skeleton

      • skull

      • ribs

      • sternum

      • vertebral column

        • cervical (neck)

        • thoracic (upper back/chest)

        • lumbar (lower back)

        • sacral (hip)

        • caudal (tail) (coccyx)

        • supports body, protection/support of nerve cord, flexibility

    • appendicular skeleton

      • forelimb bones

        • humerus (upper arm)

        • radius and ulna (lower arm)

        • carpals (wrist)

        • metacarpals (hand)

        • phalanges (fingers)

      • hindlimb bones

        • femur (thigh)

        • patella (kneecap)

        • tibia and fibula (lower leg)

        • tarsals (ankle)

        • metatarsals (foot)

        • phalanges (toes)

  • muscles

    • vertebrates have three types of muscles, classified according to structure, function and control mechanisms

      • cardiac - found only in heart

        • striated

        • involuntary

        • fibers of elongated tubes

        • possibly branched

        • multinucleated

      • smooth - surrounds and forms part of the lining of hollow organs and tubes

        • involuntary

        • ONE nucleus

      • skeletal - directly involved in locomotion

        • striated

        • voluntary

        • multinucleated

    • skeletal muscle

      • grouping of muscle fibers bound together in bundles (fascicles) by connective tissues

      • usually linked to bones by bundles of collagen fibers called tendons (dense connective tissue)

      • bones moves as muscle shortens

        • contracting muscle exerts only a pulling force - attach to bone, pulled toward or away from each other

        • requires ATP (from cell respiration)

      • example

        • muscle —> muscle fascicle —> skeletal muscle cells —> myofibrils —> thick and thin filaments (myofilaments)

        • thin filaments = actin

        • thick filaments = myosin

    • sarcomere

      • functional unit of skeletal muscle cells

      • distance from one z-line to the next

      • shortens during contraction

      • overlapping of thick and thin filaments produces striations

      • contraction

        • contracted muscle cell

        • myosin heads attach to actin filaments and pull together; shortening of sarcomeres=

        • *energy required - high energy bond of 3rd phosphate of ATP

        • adenosine P-P=P, where = is high energy bond, applied for sarcomere contraction

    • muscle cells

      • contains many mitochondria

        • powerhouse of cell

        • site of bulk ATP formation

        • requires oxygen for aerobic cellular respiration