Biology: An Introduction to Biology
Principles of Biology and Levels of Organization
- Biology defined: The study of life.
- Investigations of living things can lead to discoveries with far-reaching benefits.
- Example: Salicylic acid (aspirin) from the willow tree.
- Example: Blood pressure medicine (ACE inhibitor) from poisonous snakes.
- The study of life has revealed a set of unifying principles.
- Principles 1 through 6 define the basic features of life.
- Principle 7 and beyond are also important in all fields of biology.
Unifying Principles of Biology
- Principle 1: Cells are the simplest units of life.
- The cell theory is a foundation of biology.
- All organisms are composed of one or more cells.
- Cells are the smallest units of life.
- New cells come from pre-existing cells by cell division.
- Principle 2: Living organisms use energy.
- The maintenance of organization requires energy.
- Principle 3: Living organisms interact with their environment.
- Includes both living (e.g., predator) and non-living components (e.g., light).
- Principle 4: Living organisms maintain homeostasis.
- The amount of variability for a parameter may differ between species (e.g., body temperature in a mammal versus a reptile).
- Principle 5: The genetic material (DNA) provides a blueprint that allows organisms to grow, develop, and reproduce.
- Principle 6: Populations of organisms evolve from one generation to the next and are related by an evolutionary history.
- All organisms contain genetic material composed of DNA (deoxyribonucleic acid).
- The genetic information of a population can change over time.
- DNA sequences, the fossil record, and other lines of evidence document evolution.
- Principle 7: Structure determines function.
- Principle 8: New properties of life emerge from complex interactions.
- Principle 9: Biology is an experimental science.
- Principle 10: Biology is a quantitative science.
- Principle 11: Biologists use models and simulations to test experimental predictions and convey their ideas.
- A model is a conceptual, mathematical, or physical depiction of a real-world phenomenon.
- Principle 12: Biology affects our society.
Levels of Biological Organization
- The organization of living organisms can be analyzed at different levels of biological complexity, ranging from atoms to the biosphere:
- Atoms
- Molecules
- Cells
- Tissues
- Organs
- Organ systems
- Organism
- Population
- Community
- Ecosystem
- Biosphere
Biological Evolution
- Unity and diversity are terms often used to describe life.
- Unity refers to the common characteristics displayed by all forms of life (Principles 1-6).
- Diversity refers to the many different forms of unicellular and multicellular life.
- Evolution: The changing genetic composition of a population over time; the underlying factor that explains the unity and diversity of modern species.
Evolutionary History
- Life began on Earth as primitive cells between 3.5 to 4 billion years ago (bya).
- Evolutionary history helps us understand the structure and function of an organism.
- Evolutionary change involves modifications of pre-existing characteristics; structures may be modified to serve new purposes.
- Example: Walking limbs were modified into a dolphin’s flipper or a bat’s wing.
Mechanisms of Evolutionary Change
- Evolutionary change occurs by two mechanisms: vertical descent with mutation and horizontal gene transfer.
Vertical Descent with Mutation
- Involves transfer of genetic information from parents to offspring.
- A progression of changes within a lineage can be documented.
- New species evolve from pre-existing species by the accumulation of mutations.
- Natural selection drives an increased frequency of beneficial mutations in a population over time.
Horizontal Gene Transfer
- An organism incorporates genetic material from another organism without being the offspring of that organism; this is a distinct process of exchanging genetic information that is relatively rare.
- Can occur between different species.
- Example: Genes that confer antibiotic resistance are sometimes transferred between different species of bacteria.
Natural and Artificial Selection
- Evolution via natural selection results in adaptations that favor reproductive success.
- Adaptation: A characteristic in a species that is the result of natural selection; adaptations affect survival and reproduction.
- Artificial selection: A human-driven form of selection (e.g., human choices of desirable traits led to various breeds of dogs).
Tuskless Elephants
- The frequency of tuskless elephants appears to be increasing due to poaching.
- The tuskless condition is an adaptation that may lessen poaching.
- Less than 2% of elephants in well-protected populations are tuskless.
- Approximately 50% of the survivors of a poached population were tuskless.
- All tuskless elephants are female (females normally have tusks).
- The tuskless gene is found on the X chromosome; female elephants have two X chromosomes (XX) whereas males have only one (XY).
- Tuskless (T) is a dominant allele.
- Female tuskless elephants have the genotype X^TX^t.
- Males cannot be tuskless because a single T allele is lethal.
Classification of Living Things (Taxonomy)
- Taxonomy: The grouping of species based on common ancestry.
- Classification involves sorting at multiple levels, where species are placed into progressively smaller groups that are more closely related to each other evolutionarily.
Domains of Life
- The largest groups are the three domains of life:
- Bacteria (unicellular prokaryotes)
- Archaea (unicellular prokaryotes)
- Eukarya (unicellular and multicellular eukaryotes)
- The smallest group is the species; each species has a unique scientific name.
- Genus name is capitalized; species descriptor (specific epithet) is not capitalized.
- Both names are italicized (e.g., Amphiprion ocellaris, commonly known as the Ocellaris clownfish).
Taxonomic Hierarchy Example
- The example given for Amphiprion ocellaris is:
- Domain: Eukarya
- Supergroup: Opisthokonta
- Kingdom: Animalia
- Phylum: Chordata
- Class: Actinopterygii
- Order: Perciformes
- Family: Pomacentridae
- Genus: Amphiprion
- Species: ocellaris
Biology as a Scientific Discipline
- Science is the observation, identification, experimental investigation, and theoretical explanation of natural phenomena.
- Many scientists utilize model organisms in their research and use a scientific method to test hypotheses.
- Different branches of biology study life at different levels using a variety of tools.
- Examples: Ecology, anatomy, physiology, cell biology, molecular biology, and systems biology.
- As new tools become available, they allow scientists to ask new questions.
Levels of Investigation in Biology
- Ecologists study species in their native environments.
- Cell biologists often use microscopes to learn how cells function.
- Molecular biologists and biochemists study the molecules and macromolecules that make up cells.
- Anatomists and physiologists study how the structures of organisms are related to their functions.
- Systems biologists may study groups of molecules.
Hypotheses and Theories
- A hypothesis is a proposed explanation for a natural phenomenon.
- Based on previous observations or experiments.
- Must yield predictions that can be shown to be correct or incorrect (must be testable and falsifiable).
- Additional observations or experiments can support or reject a hypothesis, but a hypothesis is never really proven.
- Observation: Maple trees lose their leaves during autumn.
- Hypothesis: Maple trees drop their leaves in autumn because of shortened hours of sunlight.
- Alternative hypothesis: Maple trees drop their leaves in autumn because of colder temperatures.
- A theory is a broad explanation of some aspect of the natural world that is substantiated by a large body of evidence.
- Allows us to make many predictions.
- Biological theories incorporate observations, hypothesis testing, and the laws of other disciplines (physics and chemistry).
- Theories are viewed as knowledge.
- Two key attributes of a theory:
- Consistent with a vast amount of known data.
- Able to make many correct predictions.
- Example: DNA is the genetic material.
- An overwhelming body of evidence supports this theory.
Scientific Approaches
- Biologists do not follow a rigid path to discovery; rather they:
- Ask questions
- Make observations
- Conduct experiments
- Ask modified questions
- Learn from failure (lots of “trouble-shooting”)
- Repeat experiments
- Analyze data
- Try new experimental approaches
- Communicate their findings, and more…
- Researchers typically utilize 2 general approaches: discovery-based science and hypothesis testing.
Discovery-Based Science
- Involves the collection and analysis of data without having a preconceived hypothesis.
- Goal is to gather information.
- Example: Investigating a newly discovered gene without already knowing the function.
- Example: Testing drugs to look for action against disease.
- Discovery-based science often leads to hypothesis testing.
Hypothesis Testing (Scientific Method)
- Designed to be an objective way to gather knowledge.
- The steps involved:
- Observations are made regarding natural phenomena.
- These observations lead to a testable hypothesis that tries to explain the phenomena.
- Experiments are conducted to determine if the predictions are correct.
- The data are analyzed (involves use of statistical analysis).
- The hypothesis is supported or rejected based on the data.
- Data are often collected in parallel control and experimental groups.
- Groups differ by a single factor.
- Prediction: Exposure of trees to shorter amounts of daylight will cause leaves to fall.
Models in Biology
- A model is a conceptual, mathematical, or physical depiction of a real-world phenomenon.
- Biologists use models to convey their ideas, evaluate experiments, and make predictions that apply to research studies.
- Models are evaluated by their consistency with experimental data.
- Models take many forms, including:
- Structural models
- Mechanistic models
- Mathematical models
- Temporal models
- Hierarchical models
Biology as a Social Discipline
- In addition to being a scientific discipline, biology is also a social discipline.
- Biologists engage in a variety of interpersonal interactions and communication practices.
- Within a research laboratory, undergraduate students, graduate students, postdocs, technicians, and the Principal Investigator (PI) all work together.
- Different labs collaborate on projects.
- Research papers are peer-reviewed.
- At meetings, scientists discuss new data – and debate!
- You can discuss science without having “all the answers.”