Intro to Birth Defects

Learning Objectives

  1. Define and distinguish major categories of congenital anomalies

  2. Explain why timing, dose, susceptibility, and mechanism influence teratogenic outcomes

  3. Use embryologic reasoning to connect a disrupted developmental process to an anatomic defect


If a structure is abnormal at birth, what do we need to know to explain it embryologically?

  • One can trace back a birth abnormality by first comparing it to what anatomy normally forms.Then we can determine what tissue or cell population was involved in producing this structure, followed by what and when the processes they underwent in occurred. Once this context is established, a conclusion can be drawn for what processes may have failed and what would have been the consequence of those failed processes.


Fetal interruption syndrome

  • Zika virus - south and central america

  • German measles


What percent of zygotes do not implant by week 2?

  • 30-50%


What percent of recognized pregnancies end in miscarriage during the first 12 weeks?

  • 15%; most likely higher due to many unrecognized pregnancies


60% of miscarried conceptuses have what?

  • chromosomal abnormalities


How does the high rate of chromosomal abnormalities (60%) in early miscarriages contrast or non-implantation (30-50%) of zygotes by week two contrast with the causes of birth defects often seen in live births?

  • Severe genetic errors largely contribute to miscarriage and non-implantation, and function as a built in biological filter for non viable conceptuses. Live births with birth defects are more often driven by gene-environment interactions, environmental teratogens, or less severe genetic defects, like Trisomy 21 which allows for survival to term.


What percent of neonatal deaths are attributed to congenital malformations?

  • 20%


What percent of live birth have at least one birth defect?

  • 1.5-3.5% of live births


What additional percent of birth defects are discovered by end of 1st year of life? Why might this be?

  • 3-7%. Defects could be less severe or not apparent without more time such with developmental delay both physically and cognitively. Internal structural abnormalities may be asymptomatic at birth as well.


Birth defects are attributed to what?

  • Environmental exposures (5-10%)

  • Discernible genetic cause (15-20%)

  • Gene-environment interaction (70-80%) — Majority


Are development and growth interchangeable?

  • NO, development refers to a progression of stages, starting with an undifferentiated, primordial state to a highly organized, specialized and mature state, capable of carrying out a function. Growth, refers an increase in size either through the formation or enlargement of like tissues.


What are the two main types of normal morphogenesis? Example?

  • Plasia refers to cell proliferation, or an increase in the relative number of cells through mitosis. Trophy is an increase in the relative size of cells. Both processes can be seen in the development of long bones. The growth plate contains a zone of proliferation, which continuously supplies cells to the zone of cell maturation and hypertrophy, where cells enlarge. Both work together to lengthen the long bone.


What are the two main types of dysmorphogensis?

  • Dysmorphogensis can be caused by dysplasia, or an abnormal number of cells and dystrophy, abnormal cell size. Abnormally high and low cell division is referred to as hyperplasia and hypoplasia respectively. Similarly, abnormally large or small cell size is referred to as hypertrophy and hypotrophy.


What might contribute to some major birth defects having higher or lower report prevalence?

  • Many major identified birth defects affect vital organs, particularly cardiovascular defects. Oral facial defects, such as cleft palates, are especially easy to identify as they are externally presented. Clubfoot is also highly reported. Many chromosomal abnormalities are also more likely to be identified as they are often characterized by widely recognized physical features, such as trisomy 21 and due to routine prenatal screening. Birth defects that require more extensive genetic screening or internal scans may be less likely to be immediately reported if symptoms are not apparent.


What is a congenital anomaly?

  • An abnormality presented at birth in either structure, function, metabolism or behavior. It is not acquired later in life, from injury or disease.


What is Teratology?

  • The study of the causes, development and mechanism of congenital malformation or birth defects


What is dysmorphology?

  • The study of abnormal form, especially patterns of congenital anomalies


What are the mechanism by which birth defects occur?

  • malformation

  • disruption

  • deformation


What is malformation? Examples?

  • A primary error in formation of a structure. It occurs early in development (embryonic period) because the error arises as the structures begin to form.

  • Trisomy 21, orofacial cleft, anacephaly


What is disruption? Example?

  • The destruction or interference with a structure that had begun to form normally. Inherently requires an initial stage of normal development, so typically occurs later in development. Can occur in both embryonic or fetal stage.

  • Hemifacial microsomia, facial tissues were destroyed during development because the artery of the second arch failed to develop so structures didn’t receive proper blood supply


What is deformation? Example

  • An abnormal shape or position caused by mechanical forces. It requires the structure to be first formed, so it also occurs later in development. It is the only mechanism capable of self correction. It can be reversed or go away.

  • Hydrocephalus (water on the brain), deforms the skull bones because of increased intracranial pressure


What is dysplasia?

  • The production of an abnormal number of cells (high or low). This can occur during any stage of development exhibiting mitosis.


What are the patterns of multiple anomalies?

  • They can either be a sequence, syndrome or association


What is a sequence? Example?

  • Refers to when a single developmental defect results in a chain of variably expressed secondary defects that all trace back to the initiating developmental event. (domino effect). The entire cascade of events is typically well understood in a sequence.

  • The primary defect in the Pierre Robin sequence is mandibular hypolasia —> causes posterior displacement of tongue —> impaired airway —> precludes closure of palatal arches


What is a syndrome?

  • A recognizable pattern of anatomically unrelated anomalies with a shared underlying (often genetic) cause. They can contain multiple variably expressed malformations and/or sequences. This means that a given anomaly can be only partially expressed or absent between different individuals of a syndrome. It is rare for a single malformation to be unique to and always be expressed in a syndrome. The relationships of the group of anomalies is frequently not understood.


A sequence may have ____ know causes, whereas a syndrome can often be traced to ___

  • multiple; a single genetic malformation


One or more ___ may be associated with various ____

  • sequences; syndromes

  • ex. Pierre Robin sequence is often seen in stickler syndrome


What is an association?

  • A nonrandom pattern of anomalies that occurs together more often than expected, without a single established cause.


What pattern of multiple anomalies does Trisomy 21 follow?

  • Trisomy 21 is a syndrome that occurs when there is an additional third 21st chromosome. It is characterized by a recognizable pattern of anomalies such as atrioventricular septal defect, duodenal atresia, hypotonia, loose joints, flatted nose brides, upslanting palpebral, small mouth, or protruding tongue. This is a syndrome because there is one underlying cause, being a genetic defect that results in a collection of unrelated anomalies. There is no cascade of effects, rather they occur independent of one another. Different individuals with down syndrome can exhibit different anomalies and of varying degree.


Describe the Potter sequence

  • Bilateral renal agenesis, a defect in kidney development, is the underlying primary anomaly that produces a sequence of anomalies, starting with a cessation in fetal urine production. This leads to oligohydramnios, or severely low amniotic fluid, which acts as the single intermediate cause that triggers the cascade. Secondary consequential anomalies include pulmonary hypoplasia, Potter facies, and limb deformities (club foot and joint contracture), all due to the compression from the lack of space in the amniotic sac.


List the common etiology for birth defects

  • genetic factors

  • environmental factors/teratogens

  • multi factorial causes

  • mechanical factors

  • unknown causes (only 20% of cases have definite cause, 80% unknown)


How does genetics cause of birth defects?

  • chromosomal abnormalities

  • single gene disorders (gain of function, loss of function)

  • copy number or dosage effects

  • altered gene regulation (epigenetic)


How does the environment cause birth defects

  • infectious agents (fetal interruption brain syndrome —> Zika virus, rubella, German measles)

  • drugs/medications (prescription)

  • alcohol and other substances (fetal alcohols syndrome)

  • maternal diseases

  • nutrional factors (poor nutrition)

  • physical factors, radiation or mechanical constraints


What does it mean for the cause of birth defects to be multi factorial?

  • Many defects reflect both genetic susceptibility and environmental exposure. Gene-environment interactions refers to when an environmental exposure triggers an anomaly in an individual with a genetic susceptibility.

  • The same exposure may not affect every embryo in the same way

  • For example, a whole family can have a genetic mutation, but only some can present with an anomaly or birth defect. This is due to a difference in the time period where they were developing, such as an environmental effect that increase penetrance


What is penetrance?

  • An observable defect from a gene. For example, if there are two siblings, and only one has cleft palate = 50% penetrance

  • Having the cause and showing the effect

  • type of statistics


What example did we talk about in class about why timing matters?

  • Alcohol exposure during pregnancy can result in fetal alcohol syndrome. Symptom include short height, low body weight, ataxia, cognitive delays, behavioral problems, and typical facies (low nasal bridge, short nose, thin upper lip, small midface, flat philtrum ,small head, small eye openings, skin folds at corner of eye). The incidence rates are 0.2-9 in 1000 and have not significantly changed since first identified in the 1960s. Although the risks of drinking during pregnancy are well understood, the rates are still so high because recognition of pregnancy does not always immediately occur. The beginning of the third week of gestation is when a missed menstrual period may be recognized, however implantation bleeding could be mistaken for a cycle and many individuals have irregular cycles. Thus before the chance to recognize the second missed period, another four more weeks of development could have occurred. Major embryonic development occurs between weeks 3-8 and this stage has the highest sensitivity to teratogens, including alcohol. Unless an individual is actively family planning, a pregnancy could be easily missed, putting the embryo at risk of various exposures at its highest state of vulnerability.



What developmental vulnerabilities may occur in weeks 1-2, 3-8, or 8-birth of human development?

  • In the first two weeks of development, there is low sensitivity to teratogens due to the lack of an established blood supply with the mother. Rather this is an “all-or-none” stage, that may result in failure to implant or death, commonly due to severe genetic abnormalities.


What developmental vulnerabilities may occur in weeks 3-8 of human development?

  • Weeks 3-8 are the most susceptible to abnormal development and has a high sensitivity to teratogens as it is characterized organogensies, where the basic body plan and vital organs are being established. This period is when major structural malformations of the embryo is most common. It is so vulnerable due to the simultaneous rapid cell behaviors occurring including proliferation, differentiation, migration, folding, fusion, septation, and remodeling.


What developmental vulnerabilities may occur in weeks 8-birth of human development?

  • Weeks 8 and on mainly involves the growth, maturation and further functional refinement of vital organs and the fetus. Structural abnormalities are less common but functional disturbances may still occur, especially in the brain. No stage of development is safe from developmental disturbances.


____ ____ reflect disrupted developmental processes. examples?

  • structural defects

    • failed fusion → clefting/neural tube defects

    • abnormal septation → cardiovascular defects

    • altered migration/patterning → craniofacial or neural crest-related defects


A teratogenic exposure usually causes a defect only if the affected structure is ____

  • forming during that critical window


What are the principles of teratology?

  • Genotype matters

    • Embryonic and maternal genetic background can influence genetic susceptibility.

    • Is there a gene that causes this?

  • Developmental stage matters

    • The effect depends on what is forming at the time

    • During what window does this structure form?

  • Dose and duration matter

    • Greater dose or longer exposure often increases risk

    • What were you exposed to during this time? When did exposure stop?

  • Mechanism matters

    • Teratogens act by altering specific cellular or molecular processes

    • If we can identify the teratogen, which specific mechanism did it affect?

  • Outcome depends on the affected developmental process

    • proliferation, migration, fusion, septation, apoptosis, and patterning can all be disrupted


List the common teratogens

  1. alcohol

  2. retinoids

  3. antiseziure drugs

  4. classic drug teratogen

  5. infections

  6. maternal disease

  7. tobacco/nicotine

  8. radiation

  9. nutrition

  10. other medication


Alcohol

  • ethanol

  • FASD; growth, CNS, craniofacial and behavioral effects


Retinoids

  • isotretinoin, retinoic acid (vitamin A), acutane (sexually active pimply teens)

  • patterning defects, craniofacial anomalies


Antiseizure drugs

  • valproic acid

  • neural tube defect risk


Classic drug teratogen

  • thalidomide

  • limb malformations


Infections

  • rubella, CMV, toxoplasmosis, Zika

  • tissue disruption; CNS and systemic anomalies


Maternal disease

  • poorly controlled diabetes

  • increased risk of congenital anomalies


Tobacco/nicotine

  • smoking, nicotine exposure

  • growth restriction, pregnancy complications


Radiation

  • ionizing radiation

  • DNA damage, effects on rapidly dividing tissues


Nutrition

  • folate deficiency (very effective modern intervention, supplement food with folate)

  • neural tube defect risk


Other medications

  • warfarin, methotrexate

  • drug specific developmental toxicity


What are processes that can be disrupted?

  • proliferation

  • migration

  • fusion

  • septation

  • patterning

  • apoptosis/remodeling

  • differentiation

  • folding


How can proliferation be disrupted?

  • too few cells or too many cells; hypoplasia, overgrowth, agenesis


How can migration be disrupted?

  • cells fail to reach destination; neural crest or neuronal migration problems


How can fusion be disrupted?

  • persistent separation; clefts, neural tube closure defects


How can septation be disrupted?

  • persistent communication between spaces; cardiac/outflow/cloacal examples


How can patterning be disrupted?

  • wrong position or identity; axis or limb pattern abnormalities


How can apoptosis/remodeling be disrupted?

  • persistent tissue, abnormal sculpting, failure to remove temporary structures


How can differentiation be disrupted?

  • Cells fail to specialize appropriately; abnormal tissue identity of function


How can folding be disrupted?

  • Body wall, gut tube, neural tube, or organ shaped defects


How can prenatal defects be diagnosed?

  • imaging → can inform surgical planning, psychological preparation and future treatments

  • genetic screenings

    • may require analysis of fetal or placental genetic material (fluid or fetal tissue)


How does family planning help with defect prevention?

  • parents can undergo genetic screening

  • discuss risk behaviors (smoking, drinking, prescription medication)

  • maternal health optimization

  • avoid known teratogens and harmful exposures

  • prenatal vitamins, folic acid

  • vaccination


Does prenatal diagnosis replace embryological reasoning?

  • No, it identifies the defect but still requires tracing back to cause

  • Especially important to determine if subsequent pregnancies will also be at risk


Understanding birth defects requires connecting ___…

  • developmental timing → disrupted mechanism → anatomical outcome → clinical care


What is involved in care planning?

  • Anticipate related anatomy, function, delivery needs, surgical care, and long term follow up


Clinical application

  • prevention

  • prenatal detection

  • diagnosis and counseling

  • care planning


T/F birth defects are rare

  • F, birth defects affect 1/33 babies each year in the US

  • every 4.5 minutes a defect baby is born


Recessed mandible condition: name, clinical presentation, pathogenesis, incidence, diagnosis, where else is it seen?

  1. Migrognathia

  2. Underdeveloped mandible (small jaw). May cause feeding difficulties (especially in neonates). Airway compromise: risk of obstructive sleep apnea. Can sometimes “self correct” in non-syndrome cases

  3. Pathogenesis varies including genetics, reduce condyle growth or altered rotations

  4. Common in neonates

  5. Diagnosed through comorbidities (feeding, airway, growth) and jaw index

  6. Found in other craniofacial sequence and syndromes → Pierre robin, FAS, VCFS, Goldenhar


Pierre Robin Sequence: clinical definition, incidence, sequence of events, associated anomalies

  1. micrognathia, glossoptosis, airway obstruction → needs all three to diagnose

  2. ~1/8500 - 14,000 live births. Comprised of syndromic and non syndromic cases

  3. Small or slow growing jaw → posterior displacement of tongue → airway obstruction

  4. failure to thrive, feeding problems, clef palate (u-shaped)


Glossoptosis: what is it, what can it lead to, clinical criteria, causes

  1. defined as displacement of the tongue into the oropharynx

  2. Leads to upper airway obstruction which can cause

    1. feeding problems

    2. apnea

    3. cyanosis

    4. can be life threatening

    5. tends to be worse when sleeping (episodic)

  3. Clinical criteria in diagnosis

    1. Laberge 1: airway ok even supine but gets partially obstructed during feeding - mild/manageable

    2. Laberge 2: obstructed when supine but ok with prone position - positional management

    3. Laberge 3: can’t keep airway open even in prone position - these babies need intervention for sure (intubation/surgery)

  4. Causes → PRS, hypotonia (poor muscle formation)


Describe cleft palate in PRS

  • PRS patients often present with cleft palate

  • pathogenesis often related to the sequence (tongue displacement posteriorly)

  • defect is often U shaped (as opposed to v)

  • not diagnostic of PRS


Causes of PRS - developmental

  • Meckel’s Cartilage and Mandibular Outgrowth (SOX9-related pathways)

  • Hypotonia reduces mandibular movement and tongue position


Causes of PRS - Deformation/Disruption

  • Twinning/intrauterine crowding


Causes of PRS - Teratogens

  • Alcohol (FAS)

  • Antiepileptics (hydantoin, trimethadione)


Causes of PRS - Genetics

  • Deletions (Chromosomal locations on 2,4,11 and 16)

  • Duplications (Chromosomal locations on 1)

  • Translocations (Unbalanced and Balanced)

  • Mutations


Causes of PRS - Syndromic

  • Stickler (Collagens)

  • VCFS (TBX1)


Steps of developmental reasoning in birth defects

  1. what is abnormal → identify the structure or pattern

  2. what normally should have happened → describe normal development first

  3. when does that process occur → link timing to vulnerability

  4. what developmental process was disrupted → fusion, migration, septation, proliferation, remodeling etc.

  5. what caused or contributed to the disruption → genetic, environmental, multi factorial, mechanical or unknown

  6. what additional anatomy might be affected → think by tissue origin, field, sequence, or syndrome