Setting the Stage
⢠All living beings have all of the following characteristics; if even one is missing, the material is not alive:
āMetabolism involving the exchange of energy and matter with the environment
āNonrandom organization
āGrowth
āA system of heredity and reproduction
āA capacity to respond to the environment such that metabolism is not adversely affected
What is botany?
⢠Biology is the science of life or living matter in all its forms and phenomena, especially with reference to origin, growth, reproduction, structure, and behavior. (Dictionary.com)
⢠Botany is a subdiscipline of biology and is commonly defined as the scientific study of plants.
What is a plant?
⢠Most have green leaves, stems, roots, and flowers, though there are exceptions.
- Conifers (gymnosperms) donāt produce flowers.
- Mosses lack true leaves, stems, and roots and neither mosses or ferns produce flowers.
⢠Fungi were once considered plants but are now excluded.
⢠Green algae are problematic.
- Similar to plants at the cellular and biochemical level
- But share more characteristics with other algae
What do we study plants? Why does it matter?
⢠Plants changed the climate of Earth in ways that we can now live on it.
⢠Plants also produce the oxygen we breathe and the food we eat.
⢠We get cloth, paper, lumber, medicines and chemicals from plants, and plants are important to us for their aesthetic
qualities.
Using Concepts to Understand Plants
There are a few fundamental concepts that will make plant biology more easily understood.
1.Plant metabolism is based on the principles of chemistry and physics.
ā All the principles you learn in your chemistry or physics classes are completely valid for plants.
2. Plants must have a means of storing and using information.
ā Genes are the primary means of storing this information.
3. Plants reproduce, passing their genes and information on to their descendants.
ā Seeds contain information from their parents.
4. Genes, and the information they contain, can change.
ā Plants copy genes during reproduction.
ā These changes cause differences in offspring.
ā Over time, a gradual evolution occurs.
5. Plants must survive in their own environment.
ā They must be adapted to their environment.
ā Plants more suited to their environment reproduce more successfully and produce more offspring.
ā The presence of other organisms may be detrimental, beneficial, or neutral.
6. Plants are highly integrated organisms.
ā The structure and metabolism of one part tend to impact the rest of the plant.
ā An adaptation in one area is often balanced by changes in another.
7. An individual plant is the temporary result of the interaction of genes and environment.
ā The way a plant interacts with its environment is guided by genetic information that has slowly evolved over thousands of years.
ā The genetic pool of the species exists beyond the phenotype represented by a single specimen.
8. Plants do not have purpose or decision-making capacity.
ā Anthropomorphism and teleology should be avoided.
⢠Anthropomorphism: applying human characteristics to non-human organisms or things
⢠Teleology: the assumption that processes or structures have a purpose
Origin and Evolution of Plants
⢠Organisms were originally simple and increased in complexity through evolution by natural selection.
Natural Selection
⢠Organisms reproduce and have non-identical offspring whose features pass to more offspring.
⢠Offspring with features that are well-adapted to the environment reproduce more
⢠New features arise periodically by mutations.
⢠Natural selection determines whether new features are eliminated or passed on to future generations.
⢠Evolution by natural selection is a model consistent with:
āObservations of natural organisms
āExperiments
āTheoretical considerations
Origin of Plants
⢠Life on Earth began about 3.5 billion years ago with prokaryotes (bacteria and archaeans).
āPhotosynthesis arose 2.8 billion years ago in a cyanobacterium.
⢠Organelles evolved followed by division of labor and specialization.
āDNA became located in its own organelle, the cell nucleus.
āDevelopment of organelles was followed by division of labor and specialization.
āEukaryotes have nuclei (plants, fungi, animals, algae).
⢠Eukaryotes diversified, some gaining mitochondria and some gaining chloroplasts (endosymbiotic theory).
āThose with mitochondria evolved into protozoans, fungi, and animals.
āThose with mitochondria and chloroplasts evolved into plants and algae.
⢠All organisms fit into three large groups called domains: Bacteria, Archaea, and Eukarya.
⢠Early plants resembled green algae.
āThey lost many algal features and gained features suitable for life on land through natural selection.
⢠Not all organisms evolve at the same rate.
āFeatures that seem relatively unchanged are relictual features or plesiomorphic features.
⢠Flowering plants evolved 100ā120 million years ago.
āSeveral features characteristic of this group evolved at this time, including:
⢠Flowers
⢠Simple, flat, broad leaves
⢠Wood
⢠Some modern groups have derived (or apomorphic) features.
āThese are features that evolved from an ancestral feature.
āModern secondary chemical compounds are an examples of derived features.
Diversity of Plant Adaptations
⢠Over 297,000 plant species exist today; wide diversity of adaptation is important.
⢠For any aspect of the environment, many types of adaptation are possible.
āThere is no single, perfect adaptation.
āThere are alternative adaptations.
āThere are ways of coping with different environments and the multitude of factors within them.