BIOS 1700 Chapter 10 Pt. 3
Embryo Development and Cell Types
Early experiments with embryos involved manipulating cell structures intentionally.
Plunging and rolling embryos to encourage mixing of different cell types.
Observations showed a tendency for cells to associate with one another, leading to distinct layers:
Epidermal cells on the outside.
Neural cells inside, forming complex arrangements.
Cell Adhesion Molecules (CAMs)
Description of CAMs
Proteins on the cell surface that promote adhesion between cells or to the extracellular matrix (ECM).
Not just cell-to-cell interactions; also connect to structures outside the cell (ECM).
Abbreviated as ECM throughout the lecture.
Breakdown of ECM terminology:
"Extra" refers to outside.
"Matrix" refers to structures surrounding cells.
Structure of Adhesion Molecules
Three domains in adhesion proteins:
Cytoplasmic domain (interacts with the cell's interior).
Transmembrane domain (hydrophobic; spans the membrane).
Extracellular domain (interacts with other cells or ECM).
Functionality is calcium-dependent; calcium is essential for adhesion processes.
Importance of Calcium
Functions of calcium in the body
Essential for muscle contractions.
Critical for maintaining tissue integrity and other biological processes.
Common misconception explained through the necessity of calcium in various bodily functions.
Types of Cadherins
Cadherin Types
E-cadherins: specific to epithelial cells; bind exclusively to other E-cadherins.
N-cadherins: found in neuronal cells; bind only to other N-cadherins.
Cells expressing both can connect with all types, indicating flexibility in cell communication.
Cell-to-Cell and Cell-to-Extracellular Matrix Interactions
Types of adhesion interactions highlighted
Adherens Junctions: connect cells through cadherins and actin filaments.
Often described as a button-like structure for cell reinforcement.
Desmosomes: connect cells through cadherins and intermediate filaments.
Provide mechanical stability to tissues.
Hemidesmosomes: use integrins to connect cells to the ECM.
Incorporate intermediate filaments internally, anchoring the cell to the ECM.
Tight Junctions: serve as barriers to regulate passage between cells.
Gap Junctions: facilitate communication between cells via direct connections for signal exchange.
Epithelial Tissue Connections
Description of epithelial tissue
Lines external and internal surfaces in the body (e.g., skin, digestive tract, blood vessels).
Anchored to the basal lamina: a non-cellular layer rich in collagen providing structural support.
Regions of epithelial cells:
Apical region: closest to the environment.
Basolateral region: sides connecting to the basal lamina.
Epithelial cell junctions location
Tight Junctions: at the apical surface to control permeability.
Adherens Junctions: located below tight junctions; they link to actin for cell shape maintenance.
Desmosomes: provide strength through mechanical connections with adjacent cells.
Hemidesmosomes: connect cells to the basal lamina, interacting with ECM.
Gap Junctions: allow for intercellular communication, enabling direct sharing of signaling molecules.
Comparison of Plant and Animal Cell Connections
Plant cell connections
Use plasmodesmata for communication; gaps through plasma membranes between plant cells.
Middle lamella acts as a cement between cell walls, rich in carbohydrates.
Structure of cell walls:
Primary cell wall: primarily composed of cellulose.
Secondary cell wall: added for structure, influenced by age and plant type.
Differences in ECM across cell types
Plant ECM is mainly the cell wall, while animal ECM consists of material between cells.
Collagen as a common protein in animal ECM; provides strength through fibers arranged in bundles.
Health and Disease Implications
Ehlers-Danlos Syndrome: a condition related to problems with collagen structure.
Presents differently depending on the collagen type affected.
Role of integrins in cancer metastasis
Cancer cells adapt integrins to migrate through ECM, enabling metastasis to distant sites.
Specialized enzymes in cancer cells break down ECM components, facilitating movement through tissues.
Research and Future Directions
Continuing focus on ECM and cytoskeleton in various diseases, including neurodegenerative disorders like Parkinson's Disease.
Instability of cytoskeleton linked to disease progression.
Integrins not only assist in physical connections but also signal changes within cells that affect behavior.